Calceolariaceae
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Calceolariaceae is a family of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the order
Lamiales The order Lamiales (also known as the mint order) are an order in the asterid group of dicotyledonous flowering plants. It includes about 23,810 species, 1,059 genera, and is divided into about 25 families. These families include Acanthaceae, Big ...
that has been recently segregated from
Scrophulariaceae The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scr ...
. The family includes three genera, '' Calceolaria'', '' Porodittia'', and '' Jovellana'', but analysis suggests that the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
''Porodittia'' should be placed within ''Calceolaria''.Andersson, S. 2006. On the phylogeny of the genus ''Calceolaria'' (Calceolariaceae) as inferred from ITS and plastid ''matK'' sequences. ''Taxon'' 55: 125-137
abstract
.
Recent molecular phylogenies that included ''Calceolaria'' have shown not only that this genus does not belong in Scrophulariaceae (or any of the numerous families recently segregated from
Scrophulariaceae The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scr ...
) but also that it is the sister clade to the majority of the other families of the Lamiales. Morphological and chemical characters also support the separation of Calceolariaceae from Scrophulariaceae and other Lamiales. Some recent studies have supported a sister-group relationship between Calceolariaceae and
Gesneriaceae Gesneriaceae, the gesneriad family, is a family of flowering plants consisting of about 152 genera and ca. 3,540 species in the tropics and subtropics of the Old World (almost all Didymocarpoideae) and the New World (most Gesnerioideae), wi ...
. Given this close relationship, some authors opt to merge this family into Gesneriaceae as subfamily Calceolarioideae


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Disintegration of the Scrophulariaceae
Lamiales Lamiales families {{Lamiales-stub