Borogovia
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''Borogovia'' is a
troodontid Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
which lived during the Late
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
, in what is now
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
. In 1971, a Polish-Mongolian expedition discovered the remains of a small theropod at the
Altan Ula Altan may refer to: * Altan (name) * Altan (band) Altan are an Irish folk music band formed in County Donegal in 1987 by lead vocalist Mairéad Ní Mhaonaigh and her husband Frankie Kennedy. The group were primarily influenced by traditional ...
IV site, in the
Nemegt Basin The Nemegt Basin is a geographical area in the northwestern Gobi Desert, in Ömnögovi Province, southern Mongolia. It is known locally as the "Valley of the Dragons", since it is a source of many fossil finds, including dinosaurs, dinosaur eggs, ...
of Ömnögovĭ province. In 1982, upon describing '' Hulsanpes perlei'', the find was reported by Halszka Osmólska and considered by her to be a possible specimen of '' Saurornithoides''. Later she concluded that it represented a species new to science. In 1987, Osmólska named and described the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
''Borogovia gracilicrus''. The generic name is derived from the fantasy avian creatures known as 'borogoves' in the
Lewis Carroll Charles Lutwidge Dodgson (; 27 January 1832 – 14 January 1898), better known by his pen name Lewis Carroll, was an English author, poet and mathematician. His most notable works are '' Alice's Adventures in Wonderland'' (1865) and its sequ ...
poem "
Jabberwocky "Jabberwocky" is a nonsense poem written by Lewis Carroll about the killing of a creature named "the Jabberwock". It was included in his 1871 novel ''Through the Looking-Glass'', the sequel to '' Alice's Adventures in Wonderland'' (1865). The ...
", in ''
Through the Looking-Glass ''Through the Looking-Glass, and What Alice Found There'' (also known as ''Alice Through the Looking-Glass'' or simply ''Through the Looking-Glass'') is a novel published on 27 December 1871 (though indicated as 1872) by Lewis Carroll and the ...
''. The specific name is a combination of
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''gracilis'', "lightly built", and ''crus'', "shin", in reference to the elegant build of the lower leg.Osmólska, H., (1987), "''Borogovia gracilicrus'' gen. et sp. n., a new troodontid dinosaur from the Late Cretaceous of Mongolia", ''Acta Palaeontologica Polonica'' 32: 133-150 The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
specimen, ZPAL MgD-I/174, was found in the
Nemegt Formation The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, ...
, dating from the early
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the inte ...
. It consists of two lower legs of a single individual, including fragments of both tibiotarsi, the undersides of both metatarsi and the second, third and fourth toes of each foot. The tibiotarsi have an estimated length of twenty-eight centimetres. ''Borogovia'' is about two meters (6 feet) long, weighing some twenty kilograms (forty-five pounds). The tibiotarsus is very elongated. The third toe is narrow. The second phalanx of the second toe is short. The claw of the second toe is short and relatively flat. Osmólska claimed that the second toe could not be hyperextended, and suggested that it had regained a weight-bearing function, compensating for the weakness of the third toe. Andrea Cau in 2021 has called this the "falchiporan condition". ''Borogovia'' was assigned by Osmólska to the
Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...
in 1987.


See also

*
Timeline of troodontid research This timeline of troodontid research is a chronological listing of events in the history of paleontology focused on the troodontids, a group of bird-like theropod dinosaurs including animals like ''Troodon''. Troodontid remains were among the fir ...


References

{{Taxonbar, from=Q27444 Troodontids Late Cretaceous dinosaurs of Asia Fossil taxa described in 1987 Taxa named by Halszka Osmólska Nemegt fauna