Bolboceratidae
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Bolboceratidae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
. It was historically treated as a subfamily of the earth-boring dung beetles, or "dor beetles" (family
Geotrupidae Geotrupidae (from Greek γῆ ''(gē)'', earth, and τρῡπητής ''(trȳpētēs)'', borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles or dor beetles. Most excavate burrows in which t ...
), but has been considered a separate family by many authors since 1995.Bolboceratidae at TOL
/ref> Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.Smith, A.B.T. (2006) A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification. The Coleopterists Bulletin, 60:144-204.) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and
Passalidae Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles" or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their size, ranging fro ...
MCKENNA, D. D., WILD, A. L., KANDA, K., BELLAMY, C. L., BEUTEL, R. G., CATERINO, M. S., FARNUM, C. W., HAWKS, D. C., IVIE, M. A., JAMESON, M. L., LESCHEN, R. A. B., MARVALDI, A. E., MCHUGH, J. V., NEWTON, A. F., ROBERTSON, J. A., THAYER, M. K., WHITING, M. F., LAWRENCE, J. F., ŚLIPIŃSKI, A., MADDISON, D. R. and FARRELL, B. D. (2015), The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Syst Entomol, 40: 835–880. doi:10.1111/syen.12132).


Genera

* '' Athyreus'' * '' Australobolbus'' * '' Blackbolbus'' * '' Blackburnium'' * '' Bolbaffer'' * '' Bolbaffroides'' * '' Bolbelasmus'' * '' Bolbobaineus'' * '' Bolbocaffer'' * '' Bolbocerastes'' * '' Bolboceratex'' * '' Bolboceratops'' * '' Bolbocerodema'' * '' Bolboceroides'' * '' Bolbocerosoma'' * '' Bolbochromus'' * '' Bolbogonium'' * '' Bolbohamatum'' * '' Bolboleaus'' * '' Bolborhachium'' * '' Bolborhinum'' * '' Bolborhombus'' * '' Bolbothyreus'' * '' Bolbotrypes'' * '' Bradycinetulus'' * '' Cretobolbus'' * '' Elephastomus'' * '' Eubolbitus'' * '' Eucanthus'' * '' Gilletinus'' * '' Halffterobolbus'' * '' Meridiobolbus'' * '' Mesoathyreus'' * '' Mimobolbus'' * '' Namibiobolbus'' * '' Namibiotrupes'' * '' Neoathyreus'' * '' Odonteus'' * '' Parabolbapium'' * '' Parathyreus'' * '' Pereirabolbus'' * '' Prototrupes'' * '' Pseudoathyreus'' * '' Socotrabolbus'' * '' Somalobolbus'' * '' Senaspidius'' * '' Zefevazia''


Fossil genera

* †'' Amberathyreus'' Bai and Zhang 2017
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, Cenomanian * †'' Cretobolbus'' Nikolajev 1996,
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous period. It is Aptian in age and consists of sandstones, siltstones, marls and bituminous shales, deposited in a stratified lake. It is s ...
, Russia, Aptian


References


External links


Encyclopedia of Life entry
Beetle families {{Scarabaeoidea-stub