The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the

hard palate The hard palate is a thin horizontal bony plate made up of two bones of the facial skeleton, located in the roof of the mouth. The bones are the palatine process of the maxilla and the horizontal plate of palatine bone. The hard palate spans ...

in the front of the

mouth In animal anatomy, the mouth, also known as the oral cavity, or in Latin cavum oris, is the opening through which many animals take in food and issue vocal sounds. It is also the cavity lying at the upper end of the alimentary canal, bounded on ...

. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the

mandibular symphysis In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halves ...

. The mandible is the movable part of the jaw.

Structure

Sobo 1909 100 - Palatine process of maxilla

In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the

alveolar process The alveolar process () or alveolar bone is the thickened ridge of bone that contains the tooth sockets on the jaw bones (in humans, the maxilla and the mandible). The structures are covered by gums as part of the oral cavity. The synonymous ter ...

** the

palatine process In human anatomy of the mouth, the palatine process of maxilla (palatal process), is a thick, horizontal process of the maxilla. It forms the anterior three quarters of the hard palate, the horizontal plate of the palatine bone making up the re ...

* three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen

Articulations

Each maxilla articulates with nine bones: * two of the

cranium The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

: the frontal and ethmoid * seven of the face: the

nasal Nasal is an adjective referring to the nose, part of human or animal anatomy. It may also be shorthand for the following uses in combination: * With reference to the human nose: ** Nasal administration, a method of pharmaceutical drug delivery ** ...

, zygomatic, lacrimal, inferior nasal concha, palatine,

vomer The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...

, and the adjacent fused maxilla. Sometimes it articulates with the orbital surface, and sometimes with the lateral pterygoid plate of the sphenoid.

Development

The maxilla is ossified in membrane. Mall and Fawcett maintain that it is ossified from ''two'' centers only, one for the maxilla proper and one for the premaxilla. These centers appear during the sixth week of

prenatal development Prenatal development () includes the development of the embryo and of the fetus during a viviparous animal's gestation. Prenatal development starts with fertilization, in the germinal stage of embryonic development, and continues in fetal devel ...

and unite in the beginning of the third month, but the suture between the two portions persists on the palate until nearly middle life. Mall states that the frontal process is developed from both centers. The maxillary sinus appears as a shallow groove on the nasal surface of the bone about the fourth month of development, but does not reach its full size until after the second dentition. The maxilla was formerly described as ossifying from six centers, viz.: * One, the ''orbitonasal,'' forms that portion of the body of the bone which lies medial to the infraorbital canal, including the medial part of the floor of the orbit and the lateral wall of the nasal cavity. * A second, the ''zygomatic,'' gives origin to the portion which lies lateral to the infraorbital canal, including the zygomatic process. * From a third, the ''palatine,'' is developed the palatine process posterior to the incisive canal together with the adjoining part of the nasal wall. * A fourth, the ''premaxillary,'' forms the incisive bone which carries the incisor teeth and corresponds to the premaxilla of the lower vertebrates. * A fifth, the ''nasal,'' gives rise to the frontal process and the portion above the canine tooth. * And a sixth, the ''infravomerine,'' lies between the palatine and premaxillary centers and beneath the vomer; this center, together with the corresponding center of the opposite bone, separates the incisive canals from each other.

Changes by age

At birth the transverse and antero-posterior diameters of the bone are each greater than the vertical. The frontal process is well-marked and the body of the bone consists of little more than the alveolar process, the teeth sockets reaching almost to the floor of the orbit. The maxillary sinus presents the appearance of a furrow on the lateral wall of the nose. In the adult the vertical diameter is the greatest, owing to the development of the alveolar process and the increase in size of the sinus.

Function

The

of the maxillae holds the upper teeth, and is referred to as the maxillary arch. Each maxilla attaches laterally to the zygomatic bones (cheek bones). Each maxilla assists in forming the boundaries of three cavities: * the roof of the

* the floor and lateral wall of the nasal cavity * the wall of the orbit Each maxilla also enters into the formation of two fossae: the infratemporal and pterygopalatine, and two fissures, the

inferior orbital The inferior orbital fissure is formed by the sphenoid bone and the maxilla. It is located posteriorly along the boundary of the floor and lateral wall of the orbit. It transmits a number of structures, including: * the zygomatic branch of the ...

and pterygomaxillary. -When the tender bones of the upper jaw and lower nostril are severely or repetitively damaged, at any age the surrounding cartilage can begin to deteriorate just as it does after death.

Clinical significance

maxilla fracture Facial trauma, also called maxillofacial trauma, is any physical trauma to the face. Facial trauma can involve soft tissue injuries such as burns, lacerations and bruises, or fractures of the facial bones such as nasal fractures and fractur ...

is a form of

facial fracture Facial trauma, also called maxillofacial trauma, is any physical trauma to the face. Facial trauma can involve soft tissue injuries such as burns, lacerations and bruises, or fractures of the facial bones such as nasal fractures and fractur ...

. A maxilla fracture is often the result of facial trauma such as violence,

falls Falls may refer to: Places * Waterfalls or rapids * Falls, North Carolina, USA * Falls, West Virginia, USA Other uses * The ropes or wires, fed through davits, that are used to secure and lower a ship's lifeboats. * Falls (surname) * The sepa ...

or automobile accidents. Maxilla fractures are classified according to the Le Fort classification.

In other animals

Sometimes (e.g. in bony fish), the maxilla is called "upper maxilla", with the mandible being the "lower maxilla". Conversely, in birds the upper jaw is often called "upper mandible". In most vertebrates, the foremost part of the upper jaw, to which the incisors are attached in mammals consists of a separate pair of bones, the premaxillae. These fuse with the maxilla proper to form the bone found in humans, and some other mammals. In

bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...

amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...

s, and

reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...

s, both maxilla and premaxilla are relatively plate-like bones, forming only the sides of the upper jaw, and part of the face, with the premaxilla also forming the lower boundary of the nostrils. However, in mammals, the bones have curved inward, creating the palatine process and thereby also forming part of the roof of the mouth. Birds do not have a maxilla in the strict sense; the corresponding part of their beaks (mainly consisting of the premaxilla) is called "upper mandible".

Cartilaginous fish Chondrichthyes (; ) is a class that contains the cartilaginous fishes that have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fishes'', which have skeletons primarily composed of bone tissue ...

, such as sharks, also lack a true maxilla. Their upper jaw is instead formed from a cartilaginous bar that is not

homologous Homology may refer to: Sciences Biology *Homology (biology), any characteristic of biological organisms that is derived from a common ancestor *Sequence homology, biological homology between DNA, RNA, or protein sequences * Homologous chrom ...

with the bone found in other vertebrates.

Additional images

File:Maxilla anterior.png, Skull. Maxilla shown in green. Maxilla - animation 02.gif, Skull. Maxilla shown in white.

References

External links

* {{Authority control Bones of the head and neck Dental anatomy Irregular bones Human mouth anatomy