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A berry is a small, pulpy, and often edible
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries,
raspberries The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with ...
, blueberries,
blackberries The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family (biology), family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus' ...
, red currants, white currants and blackcurrants. In Britain, soft fruit is a
horticultural Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
term for such fruits. In common usage, the term "berry" differs from the scientific or botanical definition of a
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
produced from the ovary of a single flower in which the outer layer of the ovary wall develops into an edible fleshy portion (
pericarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggr ...
). The botanical definition includes many fruits that are not commonly known or referred to as berries, such as
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
s,
tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
es, cucumbers, eggplants, bananas, and
chili peppers Chili peppers (also chile, chile pepper, chilli pepper, or chilli), from Nahuatl '' chīlli'' (), are varieties of the berry-fruit of plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for t ...
. Fruits commonly considered berries but excluded by the botanical definition include strawberries, raspberries, and
blackberries The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family (biology), family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus' ...
, which are
aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
s and
mulberries ''Morus'', a genus of flowering plants in the family Moraceae, consists of diverse species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 identif ...
, which are multiple fruits.
Watermelons Watermelon (''Citrullus lanatus'') is a flowering plant species of the Cucurbitaceae family and the name of its edible fruit. A scrambling and trailing vine-like plant, it is a highly cultivated fruit worldwide, with more than 1,000 vari ...
and
pumpkins A pumpkin is a vernacular term for mature winter squash of species and varieties in the genus ''Cucurbita'' that has culinary and cultural significance but no agreed upon botanical or scientific meaning. The term ''pumpkin'' is sometimes use ...
are giant berries that fall into the category " pepos". A plant bearing berries is said to be ''bacciferous'' or ''baccate''. Berries are eaten worldwide and often used in
jam Jam is a type of fruit preserve. Jam or Jammed may also refer to: Other common meanings * A firearm malfunction * Block signals ** Radio jamming ** Radar jamming and deception ** Mobile phone jammer ** Echolocation jamming Arts and ente ...
s,
preserves Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the meth ...
, cakes, or pies. Some berries are commercially important. The berry industry varies from country to country as do types of berries cultivated or growing in the wild. Some berries such as raspberries and strawberries have been bred for hundreds of years and are distinct from their wild counterparts, while other berries, such as
lingonberries ''Vaccinium vitis-idaea'', the lingonberry, partridgeberry, mountain cranberry or cowberry, is a small evergreen shrub in the heath family Ericaceae, that bears edible fruit. It is native to boreal forest and Arctic tundra throughout the Norther ...
and cloudberries, grow almost exclusively in the wild. While many berries are edible, some are poisonous to humans, such as
deadly nightshade ''Atropa belladonna'', commonly known as belladonna or deadly nightshade, is a toxic perennial herbaceous plant in the nightshade family Solanaceae, which also includes tomatoes, potatoes, and eggplant (aubergine). It is native to Europe, North ...
and
pokeweed ''Phytolacca americana'', also known as American pokeweed, pokeweed, poke sallet, dragonberries, and inkberry, is a poisonous, herbaceous perennial plant in the pokeweed family Phytolaccaceae. This pokeweed grows . It has simple leaves on green ...
. Others, such as the white mulberry, red mulberry, and
elderberry ''Sambucus'' is a genus of flowering plants in the family Adoxaceae. The various species are commonly called elder or elderberry. The genus was formerly placed in the honeysuckle family, Caprifoliaceae, but was reclassified as Adoxaceae due to ge ...
, are poisonous when
unripe Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make the ...
, but are edible when ripe.


History

Berries have been valuable as a food source for humans since before the start of agriculture, and remain among the primary food sources of other primates. They were a seasonal staple for early hunter-gatherers for thousands of years, and wild berry gathering remains a popular activity in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
and North America today. In time, humans learned to store berries so that they could be used in the winter. They may be made into
fruit preserves Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the met ...
, and among Native Americans, mixed with meat and fats as
pemmican Pemmican (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigenou ...
. Berries also began to be cultivated in Europe and other countries. Some species of blackberries and raspberries of the genus ''
Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species. Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of the ...
'' have been cultivated since the 17th century, while smooth-skinned blueberries and cranberries of the genus ''
Vaccinium ''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (whort ...
'' have been cultivated in the United States for over a century. In Japan, between the 10th and 18th centuries, the terms and referred to many berry crops. The most widely cultivated berry of modern times, however, is the strawberry, which is produced globally at twice the amount of all other berry crops combined. The strawberry was mentioned by ancient Romans, who thought it had medicinal properties, but it was then not a staple of agriculture. Woodland strawberries began to be grown in French gardens in the 14th century. The musk strawberry (''F. moschata''), also known as the hautbois strawberry, began to be grown in European gardens in the late 16th century. Later, the Virginia strawberry was grown in Europe and the United States. The most commonly consumed strawberry, the
garden strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus ''Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
(''F. ananassa''), is an accidental hybrid of the Virginia strawberry and a Chilean variety ''
Fragaria chiloensis ''Fragaria chiloensis'', the beach strawberry, Chilean strawberry, or coastal strawberry, is one of two species of wild strawberry that were hybridized to create the modern garden strawberry (''F. × ananassa''). It is native to the Pacific Oc ...
''. It was first noted by a French gardener around the mid 18th century that, when ''F. moschata'' and ''F. virginiana'' were planted in between rows of ''F. chiloensis'', the Chilean strawberry would bear abundant and unusually large fruits. Soon after, Antoine Nicolas Duchesne began to study the breeding of strawberries and made several discoveries crucial to the science of plant breeding, such as the sexual reproduction of strawberry. Later, in the early 1800s, English breeders of strawberry made varieties of ''F. ananassa'' which were important in strawberry breeding in Europe, and hundreds of cultivars have since been produced through the breeding of strawberries.


Etymology

The Old English word berie ("berry, grape,") comes from Proto-Germanic language, Proto-Germanic *basjom (source also of Old Norse ber, Middle Dutch bere, German Beere "berry;" Old Saxon winberi, Gothic weinabasi "grape"), which is of unknown origin. This and apple are the only native fruit names.


Botanical definition

In botany, botanical terminology, a berry is a simple fruit with seeds and pulp produced from the ovary (plants), ovary of a single flower. It is fleshy throughout, except for the seeds. It does not have a special "line of weakness" along which it splits to release the seeds when ripe (i.e. it is Dehiscence (botany), indehiscent). A berry may develop from an ovary with one or more gynoecium, carpels (the female reproductive structures of a flower). The seeds are usually embedded in the fleshy interior of the ovary, but there are some non-fleshy examples such as Capsicum, peppers, with air rather than pulp around their seeds. The differences between the everyday and botanical uses of "berry" results in three categories: those fruits that are berries under both definitions; those fruits that are botanical berries but not commonly known as berries; and those parts of plants commonly known as berries that are not botanical berries, and may not even be fruits. Berries under both definitions include Blueberry, blueberries, Cranberry, cranberries, Lingonberry, lingonberries, and the fruits of many other members of the Ericaceae, heather family, as well as Gooseberry, gooseberries, Goji, goji berries and Sambucus, elderberries. The fruits of some "currants" (''Ribes'' species), such as blackcurrants, red currants and white currants, are botanical berries, and are treated as horticultural berries (or as soft fruit in the UK), even though their most commonly used names do not include the word "berry". Botanical berries not commonly known as berries include bananas,
tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
es,
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
s, eggplants (aubergines), persimmons, watermelons, and pumpkins. There are several different kinds of fruits which are commonly called berries, but are not botanical berries. Blackberry, Blackberries, Raspberry, raspberries, and Strawberry, strawberries are kinds of
aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
; they contain seeds from different ovaries of a single flower. In aggregate fruits like blackberries, the individual "fruitlets" making up the fruit can be clearly seen. The fruits of Prunus spinosa, blackthorn may be called "sloe berries", but botanically are small Drupe, stone fruits or drupes, like plums or apricots. Junipers and Taxus, yews are commonly said to have berries, but these differ from botanical berries and are instead highly modified seed-bearing Conifer cone, cones. In Juniper berry, juniper berries, used to flavour gin, the cone scales, which are hard and woody in most conifers, are instead soft and fleshy when ripe. The bright red berries of yews consist of a fleshy outgrowth (aril) almost enclosing the poisonous seed.


Cultivation

Strawberries have been grown in gardens in Europe since the 14th century. Blueberries were domesticated starting in 1911, with the first commercial crop in 1916. Huckleberries of all varieties are not fully domesticated, but domestication was attempted from 1994 to 2010 for the economically significant western huckleberry. Many other varieties of ''
Vaccinium ''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (whort ...
'' are likewise not domesticated, with some being of commercial importance.


Agricultural methods

Like most other food crops, berries are commercially grown, with both conventional pest management and integrated pest management (IPM) practices. Organic certification, Organically certified berries are becoming more widely available.:5 Many soft fruit berries require a period of temperatures between for breaking dormancy. In general, strawberries require 200–300 hours, blueberries 650–850 hours, blackberries 700 hours, raspberries 800–1700 hours, currants and gooseberries 800–1500 hours, and cranberries 2000 hours. However, too low a temperature will kill the crops: blueberries do not tolerate temperatures below , raspberries, depending on variety, may tolerate as low as , and blackberries are injured below . Spring frosts are, however, much more damaging to berry crops than low winter temperatures. Sites with moderate slopes (3–5%), facing north or east, in the northern hemisphere, near large bodies of water, which regulate spring temperature, are considered ideal in preventing spring frost injury to the new leaves and flowers. All berry crops have shallow root systems. Many land-grant university extension offices suggest that strawberries should not be planted more than five years on the same site, due to the danger of Thielaviopsis, black root rot (though many other illnesses go by the same name), which in the past has been controlled in major commercial production by annual methyl bromide fumigation but is largely prohibited now. Besides the number of years in production, soil compaction, the frequency of fumigation, and herbicide usages increase the appearance of black root rot in strawberries. Raspberries, blackberries, strawberries, and many other berries are susceptible to verticillium wilt. Blueberries and cranberries grow poorly if the clay or silt content of the soil is more than 20%, while most other berries tolerate a wide range of soil types. For most berry crops, the ideal soil is well drained sandy loam, with a pH of 6.2–6.8 and a moderate to high organic content; however, blueberries have an ideal pH of 4.2–4.8 and can be grown on muck soils, while blueberries and cranberries prefer poorer soils with lower cation exchange, lower calcium, and lower levels of phosphorus. Growing most berries organically requires the use of proper crop rotation, the right mix of cover crops, and the cultivation of the correct beneficial microorganisms in the soil. As blueberries and cranberries thrive in soils that are not hospitable to most other plants, and conventional fertilizers are toxic to them, the primary concern when growing them organically is bird management. Postharvest small fruit berries are generally stored at 90–95% relative humidity and . Cranberries, however, are frost sensitive, and should be stored at . Blueberries are the only berries that respond to ethylene, but flavor does not improve after harvest, so they require the same treatment as other berries. Removal of ethylene may reduce disease and spoilage in all berries. Precooling within one to two hours post-harvest to storage temperature, generally , via forced air cooling increases the storage life of berries by about a third. Under optimum storage conditions, raspberries and blackberries last for two to five days, strawberries 7–10 days, blueberries two to four weeks, and cranberries two to four months. Berries can be shipped under high carbon dioxide or modified atmosphere of 10–15% carbon dioxide for high carbon dioxide or 15–20% carbon dioxide and 5–10% oxygen for a modified atmosphere container to increase shelf life and prevent grey mold rot.


Breeding

Several discoveries in the science of breeding berries were made in the 18th century by Antoine Nicolas Duchesne in his work on strawberries. In the traditional technique of plant breeding, berries with specific desirable characteristics were chosen and allowed to sexually reproduce with other berries, and offspring with improved traits could then be selected and used for further crossing. Plants may be Hybrid (biology), hybridized with different species within the same genus; hybridization between different genus may also be possible, but more difficult. Breeding may seek to increase the size and yield of the fruit, improve the flavor and quality of its nutrient content, such as antioxidants, expand the harvest season, and produce cultivars with resistance to diseases, tolerance of hot or cold conditions, and other desirable traits. Advancements in molecular biology and genetic engineering allow for a more efficient and better targeted approach in the selection for a desirable genotype, via marker-assisted selection, for example. Genetically modified organism, Genetic modification techniques can also be used for breeding berries.


Horticultural soft fruit berries

Some fruit not commonly referred to as berries and not always botanically berries are included by land-grant university extension offices in their guides for berry cultivation, or in guides for identifying local wild edible and non-edible berries. Examples include Prunus maritima, beach plums, Diospyros virginiana, American persimmons, Asimina triloba, pawpaws, Malus fusca, Pacific crabapples, and Opuntia, prickly pears.


Commercial production

In the year 2005, there were of land worldwide cultivating berries, with produced.:4


Economics

In certain regions, berrypicking can be a large part of the economy, and it is becoming increasingly common for western European countries such as Sweden and Finland to import cheap labor from Thailand or Bulgaria to do the berry picking. This practice has come under scrutiny in the past years because of the low wages and poor living standard for the "berry-pickers", as well as the lack of worker safety. In the late 2010s in the US, reduced migration from Mexico and increased minimum wage standards have made finding "stoop-work" labourers to pick the strawberry crop difficult and costly.


Phytochemicals and color

Once ripening, ripened, berries have a contrasting color to their background (often of green leaves), making them visible and attractive to frugivorous animals and birds. This assists the wide biological dispersal, dispersal of the plants' seeds. Berry colors are due to natural phytochemicals, including plant pigments, such as anthocyanins, together with other flavonoids localized mainly in berry Peel (fruit), skins, seeds and leaves. Although berry pigments have antioxidant properties ''in vitro'', there is no physiological evidence established to date that berry pigments have actual antioxidant or any other functions within the human body. Consequently, it is not permitted to claim that foods containing polyphenols have antioxidant health value on product labels in the United States or Europe.


Culinary significance


Use in baked goods

Berries are commonly used in pies or tarts, such as blueberry pie, blackberry pie, and strawberry pie. Berries are often used in baking, such as blueberry muffins, blackberry muffins, berry cobblers, berry crisps, berry cakes, berry buckles, berry crumb cakes, berry tea cakes, and berry cookies. Berries are commonly incorporated whole into the batter for baking, and care is often taken so as to not burst the berries. Frozen or dried berries may be preferable for some baked berry products. Fresh berries are also often incorporated into baked berry desserts, sometimes with cream, either as a filling to the dessert or as a topping.


Beverages

Berries are often added to water and/or juiced, as in cranberry juice, which accounts for 95% of cranberry crop usage, blueberry juice, raspberry juice, goji berry juice, acai juice, aronia berry juice, and strawberry juice. Wine is the principal fermented beverage made from berries (grapes). Fruit wines are commonly made out of other berries. In most cases, sugars must be added to the berry juices in the process of Chaptalization to increase the alcohol content of the wine. Fruit wine#List of fruits and plants used to make fruit wine, Examples of fruit wines made from berries include: elderberry wine, strawberry wine, blueberry wine, blackberry wine, redcurrant wine, huckleberry wine, goji wine and cranberry wine. Berries are used in some beer style, styles of beer, particularly framboise (made with raspberry) and other fruit lambics.


Dried

Zante currant, Currants, raisins and sultana (grape), sultanas are examples of dried grape berries, and many other commercially important berries are available in dried form.


Fruit preserves

Berries are perishable fruits with a short shelf life, and are often preserved by dried fruit, drying, frozen food, freezing, pickling or making
fruit preserves Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the met ...
. Berries such as blackberry, blueberry, boysenberry, Lingonberry jam, lingonberry, loganberry, raspberry, and strawberry are often used in jams and jellies. In the United States, Native Americans in the United States, Native Americans were "the first to make preserves from blueberries".


Other usages

Chefs have created quick pickled soft fruit, such as blackberries, strawberries, and blueberries. Strawberries can be battered and quickly fried in a deep fryer. Sauces made from berries, such as cranberry sauce, can be frozen until hard, battered, and deep fried. Cranberry sauce is a traditional food item for Thanksgiving, and similar sauces can be made from many other berries such as blueberries, raspberries, blackberries, and huckleberries.


Cultural significance


Dyeing

Berries have been used in some cultures for dyeing. Many berries contain juices that can easily stain, affording use as a natural dye. For example, blackberries are useful for making dyes, especially when ripe berries can easily release juice to produce a Colour fastness, colorfast effect. ''Rubus'' berries, such as blackberry, raspberry, black raspberry, dewberry, loganberry, and thimbleberry all produce dye colors. These were once used by Native Americans. In Hawaii, the native raspberry called 'akala' was used to dye tapa cloth with lavender and pink hues, whereas berries from the dianella (plant), dianella lily were used for blue coloration, and berries from the Solanum americanum, black nightshade were used to produce green coloration.


Research

Berry consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. A 2016 review found that berry consumption can significantly lower body mass index, low density lipoprotein (LDL) and systolic blood pressure.


See also

* List of berries * List of inedible fruits


References


Further reading

* Bowling, B.L. (2005)
''The Berry Grower's Companion''
Timber Press.


External links


United States National Berry Crops Initiative
{{Authority control Berries, Fruit morphology Non-timber forest products