Berothidae

The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.

Systematics

The peculiar genus '' Lomamyia'' cannot be robustly assigned to any one subfamily. In addition, a considerable fossil diversity of beaded lacewings is known from the Late Jurassic onwards, containing numerous genera which are likewise basal or ''incertae sedis.''

Extant genera

*'' Asadeteva'' Aspöck & Aspöck, 1981
*'' Austroberothella'' Aspöck & Aspöck, 1985
*'' Berlekrumyia'' Aspöck & Aspöck, 1988
*'' Berotha'' Walker, 1860
*'' Berothimerobius'' Monserrat & Deretsky, 1999
*'' Cyrenoberotha'' MacLeod & Adams, 1968
*'' Isoscelipteron'' Costa, 1863
*'' Lekrugeria'' Navás, 1929
*'' Lomamyia'' Banks, 1904
*'' Manselliberotha'' Aspöck & Aspöck, 1988
*'' Naizema'' Navás, 1919
*'' Nodalla'' Navás, 1926
*'' Nosybus'' Navás, 1910
*'' Nyrma'' Navás, 1933
*'' Ormiscocerus'' Blanchard, 1851
*'' Podallea'' Navás, 1936
*'' Protobiella'' Tillyard, 1923
*'' Quasispermophorella'' Aspöck & Aspöck, 1986
*'' Spermophorella'' Tillyard, 1916
*'' Spiroberotha'' Adams, 1990
*'' Stenobiella'' Tillyard, 1916
*'' Tanzanberotha'' Aspöck & Hynd, 1995
*'' Trichoberotha'' Handschin, 1935
*'' Trichoma'' Tillyard, 1916

Extinct genera

* †'' Ansoberotha'' Yang et al. 2019 Burmese amber, Myanmar, Late Cretaceous ( Cenomanian)
* †'' Araripeberotha'' Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Early Cretaceous ( Aptian)
* †'' Banoberotha'' Whalley 1980 Lebanese amber, Early Cretaceous (Barremian)
* †'' Caririberotha'' Martins-Neto and Vulcano 1990 Crato Formation, Brazil, Aptian
*†'' Cantabroberotha'' Pérez-de la Fuente, Peñalver & Engel, 2020, Spanish amber, Early Cretaceous (Albian)
* †'' Cornoberotha'' Yang et al. 2019 Burmese amber, Myanmar, Cenomanian
* †'' Dasyberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
* †'' Elektroberotha'' Makarkin and Ohl 2015 Baltic amber, Eocene
* †'' Epimesoberotha'' Jepson et al. 2012 Lulworth Formation, United Kingdom, Early Cretaceous (Berriasian)
* †'' Ethiroberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
*†'' Haploberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
*†'' Iceloberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
*†'' Jersiberotha'' Grimaldi 2000 Burmese amber, Myanmar, Cenomanian New Jersey amber, Late Cretaceous (Turonian)
*†'' Krokhathone'' Khramov 2015 Karabastau Formation, Kazakhstan, Middle/Late Jurassic (Callovian/ Oxfordian)
*†'' Maculaberotha'' Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian
*†'' Magniberotha'' Yuan et al. 2016 Burmese amber, Myanmar, Cenomanian
*†subfamily Mesithoninae Panfilov 1980
**†'' Berothone'' Khramov 2015 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
**†'' Mesithone'' Panfilov 1980 Karabastau Formation, Kazakhstan, Callovian/Oxfordian Zaza Formation, Turga Formation, Russia, Aptian
**†'' Pseudosisyra'' Makarkin 1999 Zaza Formation, Russia, Aptian
*†'' Microberotha'' Archibald and Makarkin 2004 Hat Creek amber, Canada, Eocene (Ypresian)
*†'' Nascimberotha'' Grimaldi 2000 New Jersey amber, Turonian
*†'' Oloberotha'' Ren and Guo 1996 Yixian Formation, China, Aptian
*†'' Osmyloberotha'' Khramov 2021 Burmese amber, Myanmar, Cenomanian
*†'' Protoberotha'' Huang et al. 2019 Burmese amber, Myanmar, Cenomanian
*†'' Sibelliberotha'' Azar and Nel 2013 Lebanese amber, Barremian
*†'' Sinosmylites'' Hong 1983 Daohugou, Haifanggou Formation, China, Callovian, Karabastau Formation, Kazakhstan, Callovian/Oxfordian, Ulaan-Ereg Formation, Mongolia, Late Jurassic ( Tithonian)
*†'' Systenoberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
*†'' Telistoberotha'' Engel and Grimaldi 2008 Burmese amber, Myanmar, Cenomanian
*†'' Xiaoberotha'' Shi et al. 2019 Burmese amber, Myanmar, Cenomanian
* †'' Xenoberotha'' Makarkin 2017 Green River Formation, United States, Eocene

References

Bugguide.net. Family Berothidae - Beaded Lacewings

Mantispoidea
Taxa named by Anton Handlirsch
Neuroptera families

{{Neuroptera-stub