Banksia laevigata subsp. laevigata
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''Banksia laevigata'' subsp. ''laevigata'', the tennis ball banksia, is a subspecies of small woody
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
in the plant genus '' Banksia''. It occurs in Western Australia's semi-arid shrubland. It and the closely related ''B. laevigata'' subsp. ''fuscolutea (golden ball banksia) are the two subspecies of the species '' Banksia laevigata''.


Taxonomy

The species ''B. laevigata'' was first described by Swiss botanist Carl Meissner in 1856, after being originally collected by James Drummond in 1848. No separate subspecies were recognised until
Alex George Alexander or Alex George may refer to: *Alex George (botanist) (born 1939), Australian botanist * Alexander L. George (1920–2006), American political scientist * Alexander George (philosopher), American philosopher *Alex George (motorcyclist), Sc ...
collected specimens of subspecies '' fuscolutea'' east of Hyden in 1964, and formally named it two years later. The naming of a new subspecies automatically created the
autonym (botany) In botanical nomenclature, autonyms are automatically created names, as regulated by the ''International Code of Nomenclature for algae, fungi, and plants'' that are created for certain subdivisions of genera and species, those that include the ty ...
subspecies ''laevigata''. The common name is derived from the resemblance of its inflorescences to tennis balls. George described its nearest probable relative as '' Banksia audax'' and placed it in the series Cyrtostylis, which he concedes is rather heterogeneous. The series was split into three in the 1996 morphological cladistic analysis by botanists Kevin Thiele and
Pauline Ladiges Pauline Yvonne Ladiges (born 1948) is a botanist whose contributions have been significant both in building the field of taxonomy, ecology and historical biogeography of Australian plants, particularly Eucalypts and flora, and in science educa ...
, with ''B. laevigata'' placed in the new series Ochraceae with ''B. audax'' and ''B. benthamiana'' in their arrangement. This was reinforced in American botanist Austin Mast's 2002 analysis, as ''B. laevigata'' came out as sister to three groups; the first the pair comprising ''B. audax'' and ''B. benthamiana''. Early in 2007, Mast and Thiele rearranged the genus ''Banksia'' by merging ''Dryandra'' into it, and published ''B.'' subg. ''Spathulatae'' for the taxa having spoon-shaped cotyledons; thus ''B.'' subg. ''Banksia'' was redefined as encompassing taxa lacking spoon-shaped cotyledons. They foreshadowed publishing a full arrangement once DNA sampling of ''Dryandra'' was complete; in the meantime, if Mast and Thiele's nomenclatural changes are taken as an interim arrangement, then ''B. laevigata'' subsp. ''laevigata'' is placed in ''B.'' subg. ''Banksia''.


Description

The tennis ball banksia is a shrub to 3.5 metres (11.5 ft) high with grey, flaky bark. New growth occurs in summer, and the young stems and leaves are covered in fine hairs, which fall away with age. There are 2 variants distinguished by their leaf margins, one along the Fitzgerald River with rather obtuse leaf teeth, the other in the Ravensthorpe Ranges with larger, acute teeth similar to those of subsp. ''fuscolutea''. The former is represented by the type. The inflorescences occur in spring and are oval in shape and a greenish yellow in colour. Flowers appear from October to December.


Distribution and habitat

''Banksia laevigata'' subsp. ''laevigata'' grows in shrubland on stony soils of
spongolite {{unreferenced, date=February 2019 Spongolite is a stone made almost entirely from fossilised sponges. It is light and porous. The silica spicules fossilised with the sponges makes the material hazardous to handle by being highly abrasive. Becau ...
or
laterite Laterite is both a soil and a rock type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by ...
, along the lower
Fitzgerald River The Fitzgerald River is a river in the Great Southern region of Western Australia. Surveyor General John Septimus Roe named the river during expeditions in the area in 1848 after the governor of Western Australia of the day, Charles Fitzge ...
and in the Ravensthorpe Ranges in inland southern Western Australia. ''Banksia laevigata'' subsp. ''laevigata'' is classified as ''Priority Four - Rare'' on the Declared Rare and Priority Flora List under the Wildlife Conservation Act of Western Australia. That is, it is a taxon which has been adequately surveyed and which, while being rare (in Australia), are not currently threatened by any identifiable factors. It requires monitoring every 5–10 years.


Ecology

Like other banksias, the tennis ball banksia is likely to play host to a variety of pollinators. Ants, bees and wasps have been recorded thus far.


Cultivation

Although like many western banksias it is sensitive to '' Phytophthora cinnamomi'' dieback, it has been successfully grown on Australia's east coast. It has also been successfully grafted onto '' Banksia integrifolia''.


References

* {{Taxonbar, from=Q4856632 laevigata subsp. laevigata Eudicots of Western Australia Plant subspecies