''Balaena'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel th ...

(whale) in the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Balaenidae Balaenidae () is a family of whales of the parvorder Mysticeti that contains two living genera: the right whales (genus '' Eubalaena''), and in a separate genus, the closely related bowhead whale (genus '' Balaena''). Evolutionary history Bal ...

. ''Balaena'' is considered a

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

genus, as it has only a single

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

species, the

bowhead whale The bowhead whale (''Balaena mysticetus'') is a species of baleen whale belonging to the family Balaenidae and the only living representative of the genus '' Balaena''. They are the only baleen whale endemic to the Arctic and subarctic waters, a ...

(''B. mysticetus''). It was named in 1758 by

Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

, who at the time considered all of the

right whale Right whales are three species of large baleen whales of the genus ''Eubalaena'': the North Atlantic right whale (''E. glacialis''), the North Pacific right whale (''E. japonica'') and the Southern right whale (''E. australis''). They are c ...

s (and the bowhead) as a single species. Historically, both the family Balaenidae and genus ''Balaena'' were known by the common name, "right whales", however ''Balaena'' are now known as bowhead whales. "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species." Balaena-montalionisBologna

Throughout history, the family Balaenidae has been the subject of great taxonomic debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. In the early whaling days, they were all thought to be a single species, ''Balaena mysticetus''. Eventually, it was recognized that bowheads and right whales were in fact different. Later, morphological factors such as differences in the skull shape of northern and southern right whales indicated at least two species of right whale—one in the Northern Hemisphere, the other in the

Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smal ...

. As recently as 1998, Dale Rice, in his comprehensive and otherwise authoritative classification, ''Marine mammals of the world: systematics and distribution'', listed just two species: ''Balaena glacialis'' (the right whales) and ''Balaena mysticetus'' (the bowheads). A DNA study by Rosenbaum in 2000, and another study by Churchill in 2007 finally provided clear evidence to conclude that the ''three'' living right whale species do comprise a

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...

lineage, ''distinct from the bowhead'', and that the bowhead and the right whales are rightly classified into two separate genera. The right whales, therefore, are now officially in the genus '' Eubalaena''. The fossil record of ''Balaena'', dating to the late Miocene, encompasses ten species known from finds in Europe, North America, and South America.The Paleobiology Database ''Balaena'' entry
accessed on 21 December 2010 ''Balena'', described by Scopoli in 1777, and ''Leiobalaena'', described by Eschricht in 1849, are

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...

s of ''Balaena''.

Taxonomy

* ''Balaena''

Bowhead whale The bowhead whale (''Balaena mysticetus'') is a species of baleen whale belonging to the family Balaenidae and the only living representative of the genus '' Balaena''. They are the only baleen whale endemic to the Arctic and subarctic waters, a ...

s **

† A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...

Balaena affinis ''Balaena'' is a genus of cetacean (whale) in the family Balaenidae. ''Balaena'' is considered a monotypic genus, as it has only a single extant species, the bowhead whale (''B. mysticetus''). It was named in 1758 by Linnaeus, who at the time con ...

'' (

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Red Crag Formation, UK) ** † ''

Balaena montalionis ''Balaena'' is a genus of cetacean (whale) in the family Balaenidae. ''Balaena'' is considered a monotypic genus, as it has only a single extant species, the bowhead whale (''B. mysticetus''). It was named in 1758 by Linnaeus, who at the time con ...

'' (

Piacenzian The Piacenzian is in the international geologic time scale the upper stage or latest age of the Pliocene. It spans the time between 3.6 ± 0.005 Ma and 2.588 ± 0.005 Ma (million years ago). The Piacenzian is after the Zanclean and is followed ...

;

Casina Casina ( egl, label= Montanaro, Caṡîna ; egl, label= Reggiano, Caṡèina ) is a '' comune'' (municipality) in the Province of Reggio Emilia in the Italian region Emilia-Romagna, located about west of Bologna and about southwest of Reggio ...

, Italy). ** '' Balaena mysticetus'',

(

Late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch withi ...

Modern Modern may refer to: History *Modern history ** Early Modern period ** Late Modern period *** 18th century *** 19th century *** 20th century ** Contemporary history * Moderns, a faction of Freemasonry that existed in the 18th century Philosophy ...

) ** † ''

Balaena ricei ''Balaena'' is a genus of cetacean (whale) in the family Balaenidae. ''Balaena'' is considered a monotypic genus, as it has only a single extant species, the bowhead whale (''B. mysticetus''). It was named in 1758 by Linnaeus, who at the time con ...

'' (Piacenzian; Rice's Pit,

Yorktown Formation The Yorktown Formation is a mapped bedrock unit in the Coastal Plain of Maryland, Virginia, North Carolina and South Carolina. It is overconsolidated and highly fossiliferous. Description The Yorktown is composed largely of overconsolidated san ...

, Virginia, USA).

Dubious species

* † ''

Balaena arcuata ''Balaena'' is a genus of cetacean (whale) in the family Balaenidae. ''Balaena'' is considered a monotypic genus, as it has only a single extant species, the bowhead whale (''B. mysticetus''). It was named in 1758 by Linnaeus, who at the time con ...

'' (

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

/Pliocene; Belgium) * † '' Balaena forsythmajori'' (

Zanclean The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, a ...

; Case il Poggio. Italy) * † '' Balaena larteti'' (

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

;

Landes ''Landes'', or ''Lanas'' in Gascon, means moorland or heath. ''Landes'' and ''Lanas'' come from the Latin ''plānus'' meaning “‘flat, even, level, plain’”. They are therefore cognate with the English plain (and plane), the Spanish word ''l ...

, France) * † '' Balaena macrocephalus'' (

Langhian The Langhian is, in the ICS geologic timescale, an age or stage in the middle Miocene Epoch/Series. It spans the time between 15.97 ± 0.05 Ma and 13.65 ± 0.05 Ma (million years ago) during the Middle Miocene.GeoWhen (2007) The Langhian was ...

; Sos, France) * † '' Balaena simpsoni'' (Miocene/Pliocene; Ancud, Chile) (possibly a member of

Neobalaenidae Neobalaenidae is a family of baleen whales (suborder Mysticeti) including the extant pygmy right whale. Although traditionally considered related to balaenids, a recent phylogenetic study by Fordyce and Marx (2013) recovered the living pygmy righ ...

R. Donoso-Barros. 1975. Contribucion al conocimiento de los cetaceos vivientes y fosiles del territorio de Chile. Gayana Zoologica 36:1-127)

Species inquirenda

* † '' Balaena pampaea'' ( Calabrian; Pampeana Formation,

Bahía Blanca Bahía Blanca (; English: White Bay) is a city in the southwest of the province of Buenos Aires, Argentina, by the Atlantic Ocean, and is the seat of government of the Bahía Blanca Partido. It had 301,572 inhabitants according to the . It is th ...

, Argentina) † =

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

species

References

{{Taxonbar, from=Q4820658 Balaenidae Mammal genera Mammal genera with one living species Extant Miocene first appearances Taxa named by Carl Linnaeus