Ardipithecus kadabba
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''Ardipithecus kadabba'' is the scientific classification given to fossil remains "known only from teeth and bits and pieces of skeletal bones", originally estimated to be 5.8 to 5.2 million years old, and later revised to 5.77 to 5.54 million years old. According to the first description, these fossils are close to the common ancestor of chimps and humans. Their development lines are estimated to have parted 6.5–5.5 million years ago. It has been described as a "probable
chronospecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
" (i.e. ancestor) of '' A. ramidus''. Although originally considered a subspecies of ''A. ramidus'', in 2004 anthropologists
Yohannes Haile-Selassie Yohannes Haile-Selassie Ambaye (born 23 February 1961) is an Ethiopian paleoanthropologist. An authority on pre-''Homo sapiens'' hominids, he particularly focuses his attention on the East African Rift and Middle Awash valleys.Mangels, John (200 ...
,
Gen Suwa Gen Suwa (born 1954) is a Japanese paleoanthropologist. He is known for his contributions to the understanding of the evolution of early hominids, including the discovery of a tooth from a hominid that was more than one million years older than the ...
, and Tim D. White published an article elevating ''A. kadabba'' to species level on the basis of newly discovered teeth from
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
. These teeth show "primitive morphology and wear pattern" which demonstrate that ''A. kadabba'' is a distinct species from ''A. ramidus''. The specific name comes from the Afar word for "basal family ancestor".


Taxonomy

Fossil remains were first described in 2001 by Ethiopian paleoanthropologist
Yohannes Haile-Selassie Yohannes Haile-Selassie Ambaye (born 23 February 1961) is an Ethiopian paleoanthropologist. An authority on pre-''Homo sapiens'' hominids, he particularly focuses his attention on the East African Rift and Middle Awash valleys.Mangels, John (200 ...
based on bones collected from five localities in the
Middle Awash The Middle Awash is a paleoanthropological research area in the Afar Region along the Awash River in Ethiopia's Afar Depression. It is a unique natural laboratory for the study of human origins and evolution and a number of fossils of the earliest ...
, Ethiopia. Haile-Selassie initially classified them as ''
Ardipithecus ramidus ''Ardipithecus ramidus'' is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). ''A. ramidus'', unlike modern hominids, has adaptations for both walking on two legs ( bipedality) and life i ...
kadabba'', with ''kadabba'' deriving from the Afar language meaning " basal family ancestor". In 2004, he, along with Japanese paleoanthropologist
Gen Suwa Gen Suwa (born 1954) is a Japanese paleoanthropologist. He is known for his contributions to the understanding of the evolution of early hominids, including the discovery of a tooth from a hominid that was more than one million years older than the ...
and American paleoanthropologist Tim D. White, elevated it to species level as ''A. kadabba'' based on apparently primitive features compared to ''A. ramidus''. ''A. kadabba'' is considered to have been the direct ancestor of ''A. ramidus'', making ''Ardipithecus'' a
chronospecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
. Along with elevating it to species level, they suggested that ''Ardipithecus'', ''
Sahelanthropus ''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed '' ...
'', and ''
Orrorin ''Orrorin tugenensis'' is a postulated early species of Homininae, estimated at and discovered in 2000. It is not confirmed how ''Orrorin'' is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecin ...
'' could potentially belong to the same genus. In 2008, American paleoanthropologists Bernard Wood and Nicholas Lonerga said that the larger ape-like canines of ''A. kadabba'' cast doubt on its assignment to the human line, but the position of ''Ardipithecus'' near humans has been reaffirmed by the discoverers and colleagues. They see a lineage of apes whose teeth continually reduce in size: ''A. kadabba''–''A. ramidus''–''
Australopithecus anamensis ''Australopithecus anamensis'' is a hominin species that lived approximately between 4.2 and 3.8 million years ago and is the oldest known ''Australopithecus'' species, living during the Plio-Pleistocene era. Nearly one hundred fossil specimens ...
''–'' Au africanus'', though they are unsure if ''Ardipithecus'' were the ancestors to these ''Australopithecus'' species, or were only closely related.
Evolutionary tree A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
according to a 2019 study:


Description

''A. kadabba'' is known from nineteen specimens which reveal elements of the teeth, jaw, feet, and hands and arms. The
holotype specimen A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, ALA-VP-2/10, is a right lower jaw fragment with a third molar, discovered in December 1997, and five associated left lower jaw teeth or root fragments collected in 1999. This correction of the initial allocation of the fossil record was based on the argument that ''Ardipithecus kadabba'' had more "primitive" features than other ''Ardipithecus'' fossils. ''Ardipithecus kadabba'' thus also has a greater similarity with the genera ''Sahelanthropus'' and ''
Orrorin ''Orrorin tugenensis'' is a postulated early species of Homininae, estimated at and discovered in 2000. It is not confirmed how ''Orrorin'' is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecin ...
''. These statements were based on additional bone finds that came to light in November 2002 and were dated at 5.8 to 5.6 million years. At the same time, it was emphasized that evidence could be found of a reduced "honing" complex, traces on the teeth that arise when the canines rub against each other when biting, constantly sharpening their peaks, which has been found in all older finds. The loss of this feature in the successor species of ''Ardipithecus ramidus'' has been used for the allocation of discoveries in that line of development of great apes that led to the australopithecines and the genus ''Homo''.


Paleoecology

The first description suggested that ''Ardipithecus kadabba'' lived in a habitat that consisted of forests, wooded savannas, and open water areas, as had been described for ''
Sahelanthropus ''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed '' ...
''.


References


External links


''Ardipithecus kadabba''
on the Smithsonian Institution's Human Origins Program
Human Timeline (Interactive)
Smithsonian,
National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...
(August 2016). {{Taxonbar, from=Q134987 Pliocene primates Miocene primates of Africa Pliocene mammals of Africa Transitional fossils Hominin fossils Fossil taxa described in 2001