The Arbutoideae are a

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus Genus ( plural genera ) is a taxonomic rank used in the biological classifica ...

in the plant family

Ericaceae The Ericaceae are a family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with c.4250 known species spread across 124 genera, making it th ...

. Phylogenetic analysis supported all genera of the subfamily as

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gr ...

, except ''

Arbutus ''Arbutus'' is a genus of 12 accepted speciesAct. Bot. Mex no.99 Pátzcuaro abr. 2012.''Arbutus bicolor''/ref> of flowering plants in the family Ericaceae, native to warm temperate regions of the Mediterranean, western Europe, the Canary Islan ...

''. Moreover, it was suggested that the non-sister relationship between

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...

and North American species may be explained by a once widespread distribution in the Northern hemisphere before the

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

. The genera ''Arbutus'', ''Arctostaphylos'', ''Comarostaphylis'' form a particular type of

mycorrhiza A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plan ...

l symbiosis with the fungus, '' Arbutoid mycorrhiza'', which resembles

ectomycorrhiza An ectomycorrhiza (from Greek ἐκτός ', "outside", μύκης ', "fungus", and ῥίζα ', "root"; pl. ectomycorrhizas or ectomycorrhizae, abbreviated EcM) is a form of symbiotic relationship that occurs between a fungal symbiont, or my ...

Genera List

References

External links

*
Arbutoideae in the Flora of North America
Asterid subfamilies {{Ericaceae-stub