Anopterus glandulosus
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''Anopterus glandulosus'', commonly known as native laurel or Tasmanian laurel, is a species of shrub or small tree in the family
Escalloniaceae Escalloniaceae is a family of flowering plants consisting of about 130 species in seven genera. In the APG II system it is one of eight families in the euasterids II clade (campanulids) that are unplaced as to order. More recent research has prov ...
. Endemic to south and southwestern Tasmania, ''A. glandulosus'' is widespread in the moist understoreys of
Tasmanian temperate rainforests The Tasmanian temperate rain forests are a temperate broadleaf and mixed forests ecoregion in western Tasmania. The ecoregion is part of the Australasian realm, which includes Tasmania and Australia, New Zealand, New Guinea, New Caledonia, and ad ...
and wet
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
forests from sea level to mountainous regions below above sea level.Curtis WM & Morris DI 1993, ''The student’s flora of Tasmania'', 2nd edn St David’s Park publishing, Hobart, Tasmania.


Taxonomy

The name '' Anopterus'' comes from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
words ''ano'' (upwards) and ''pteron'' (wing), referring to its characteristic winged seeds, while ''glandulosus'' refers to the glandular tips present at the end of each leaf serration. The signature 'Labill' often appears after the Latin name of this species as it was first described by
Jacques Labillardière Jacques-Julien Houtou de Labillardière (28 October 1755 – 8 January 1834) was a French biologist noted for his descriptions of the flora of Australia. Labillardière was a member of a voyage in search of the La Pérouse expedition. He pub ...
in 1805. It is one of two Tasmanian plants in the ancient Gondwanan family
Escalloniaceae Escalloniaceae is a family of flowering plants consisting of about 130 species in seven genera. In the APG II system it is one of eight families in the euasterids II clade (campanulids) that are unplaced as to order. More recent research has prov ...
, the other plant being '' Tetracarpaea tasmanica''. '' Anopterus'' is an endemic Australian genus and ''Anopterus glandulosus'' is one of two species in the genus. The other species, '' Anopterus macleayanus'' or ‘Queensland Laurel’/ ‘Macleay Laurel’ is found in subtropical, warm and cool temperate forests near the coast in New South Wales and Queensland.


Description

''Anopterus glandulosus'' usually grows as small understorey shrub ranging from in height and in width, however, can be grow as a small canopy tree up to . In shaded understorey conditions its growth habit is often straggly with branches forming layers resulting in coppiced growth. Leaves are large, in length and in width. Leaves are thick, dark green with a glabrous surface and glossy appearance. Leaf margins are serrated with blunt tips and a black gland at the apex of each serration. Leaf shape ranges from lanceolate elliptical to oblanceolate with an acute apex and leaf base which narrows into a short petiole. Flowering occurs in late spring and often again in autumn. The flower heads, known as
inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...
s, are borne in terminal racemes which are about the same length as the leaves. Flowers are bell shaped and approximately in diameter, occurring on slender pedicels which are often recurved. The flowers are composed of six ovate petals, in length which overlap and are colored white or pink flushed. Flowers usually have six sepals, which are joined at the base with spreading lobes, and six stamens opposite the sepals, which are filamentous and shorter than petals.) Each flower has two large joined carpels which are bright green in colour. The ovary is superior and conical, tapering to a short style and bifid stigma.


Similar species

The foliage arrangement and habit of ''Anopterus glandulosus'' are similar to '' Cenarrhenes nitida'' (Port Arthur/ native plum) with opposite leaves arranged in apparent spreading whorls.Wiltshire R & Jordan 2009,“TreeFlip, Life-size guide to the trees of Tasmania”, School of Plant Science University of Tasmania & CRC for Forestry ''A. glandulosus'' can be distinguished from ''Cenarrhenes nitida'' by a tendency for leaves to occur in clusters at the ends of branches. ''Cenarrhenes nitida'' can also be distinguished by the absence of glands at leaf tips and an unpleasant foetid odour when leaves are crushed.


Ecology and distribution

''Anopterus glandulosus'' is a slow growing shrub to small tree which occurs in wet
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
forest and rainforests, in the south and west of Tasmania at elevations below 1200 m. ''A. glandulosus'' prefers cool, moist conditions in partly shaded conditions with well drained soils, from loamy to sandy and highly organic soils. More seedlings of this species germinate and grow on rotting fallen logs than soil. It most commonly occurs as an
understorey In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
shrub often under a canopy of ''
Nothofagus cunninghamii ''Nothofagus cunninghamii,'' commonly known as myrtle beech or Tasmanian myrtle, is the dominant species of cool temperate rainforests in Tasmania and Southern Victoria. It has low fire resistance and grows best in partial shade conditions. It ...
'' (myrtle beech). It is susceptible to the plant disease ''
Phytophthora cinnamomi ''Phytophthora cinnamomi'' is a soil-borne water mould that produces an infection which causes a condition in plants variously called " root rot", "dieback", or (in certain ''Castanea'' species), "ink disease". The plant pathogen is one of the wo ...
'', which causes ‘root rot’ or ’dieback’, and does not tolerate excess phosphorus. However, this species is highly adaptable and tolerates harsh conditions including to frost and snow. Due to its rainforest habitat it is also adapted for continuous regeneration and regenerates well after disturbance, often undergoing rapid regrowth to form thickets in response to mechanical damage. The attractive and strongly scented flowers produce large amounts of nectar, attracting insects and birds and birds that feed on both nectar and insects such as honeyeaters.


Status

The species was listed as not threatened and present in state reserves by the National Parks and Wildlife service Tasmania in 1997 and remains widespread and non-threatened today.Howells, C (eds), Whiting J, Roberts J, Reeves R, Tayler F & Tayler V 2015, ''Tasmania’s natural flora'', 2nd edn, Australian Plants Society Inc., Hobart Group, Hobart, Tasmania.


Cultivation

''Anopterus glandulosus'' is an attractive and easily cultivated plant, making it an excellent for gardens. Successful cultivation can be achieved by germination of fresh seed or propagation from semi-hardwood cuttings taken in March or April. Germination from seed can take 4–5 months, and requires constant moisture. This species tolerates full sun to shade and is easily grown in a semi- protected environment, against a wall or fence, and grows particularly well as a container plant. It requires ample water in summer and benefits from application of non-phosphate based organic fertilisers along with manure and mulch. This species also responds well to annular pruning, and its attractive evergreen foliage can be harvested for floriculture.


References


External links

* * {{Taxonbar, from=Q2852322 Asterids of Australia Endemic flora of Tasmania Trees of Australia Escalloniaceae Plants described in 1805