Anaxyelidae
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Anaxyelidae is a family of incense cedar wood wasps in the order
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typic ...
. It contains only one living genus, '' Syntexis'', which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jurassic, belonging to over a dozen extinct genera, with a particularly high diversity during the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145& ...
. ''Syntexis'' lay eggs in the sapwood of conifers, preferring recently burnt wood.


Genera

These genera belong to the family Anaxyelidae: Taxonomy after * Subfamily Syntexinae: ** '' Syntexis'' Rohwer, 1915 Western North America, Recent ** † '' Cretosyntexis'' Rasnitsyn & Martinez-Delclos, 2000 La Pedrera de Rubies Formation, Spain, Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
) ** † '' Curiosyntexis'' Kopylov, 2019 Ola Formation Russia, Late Cretaceous (
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...
) ** † '' Daosyntexis'' Kopylov, Rasnitsyn, Zhang & Zhang, 2020
Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China, Middle Jurassic (
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
) ** † '' Dolichosyntexis'' Kopylov, 2019
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous period. It is Aptian in age and consists of sandstones, siltstones, marls and bituminous shales, deposited in a stratified lake. It is s ...
, Russia, Early Cretaceous (
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
) ** † '' Eosyntexis'' Rasnitsyn, 1990 Purbeck Group, United Kingdom, Early Cretaceous (
Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ag ...
) La Pedrera de Rubies Formation, Spain, Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
), Turga Formation, Russia, Aptian, Spanish amber, Early Cretaceous (
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...
) ** † '' Orthosyntexis'' J. Gao, Engel, Shih, & T. Gao, 2021
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, Late Cretaceous (
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in ...
) ** † '' Parasyntexis'' Kopylov, 2019 Khasurty, Zaza Formation, Russia, Early Cretaceous (
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
) ** † '' Sclerosyntexis'' Wang, Ren, Kopylov & Gao, 2020
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, Late Cretaceous (
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in ...
) *Anaxyelinae ** † '' Anasyntexis'' Rasnitsyn, 1968 Karabastau Formation, Middle/Late Jurassic (
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
- Oxfordian) ** † '' Anaxyela'' Martynov, 1925 Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian) **† '' Brachysyntexis'' Rasnitsyn, 1969 Daohugou, China, Middle Jurassic (Callovian), Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian),
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its exq ...
, China, Aptian, Khasurty, Zaza Formation, Russia, Early Cretaceous (Aptian) ** † '' Dolichostigma'' Rasnitsyn, 1968 Zaza Formation, Russia, Early Cretaceous (Aptian) ** † '' Kempendaja'' Rasnitsyn, 1968 Khaya Formation, Russia, Late Jurassic (
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by ...
) ** † '' Kulbastavia'' Rasnitsyn, 1968 Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian) ** † '' Mangus'' Kopylov, 2019 Dzun-Bain Formation, Mongolia, Early Cretaceous (Aptian) ** † '' Sphenosyntexis'' Rasnitzyn, 1969 Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian) ** † '' Syntexyela'' Rasnitsyn, 1968 Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian), Yixian Formation, China, Aptian ** † '' Urosyntexis'' Rasnitzyn, 1969 Karabastau Formation, Middle/Late Jurassic (Callovian-Oxfordian), Khasurty, Zaza Formation, Russia, Early Cretaceous (
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
)


References


Further reading

* * Sawflies Extant Jurassic first appearances {{hymenoptera-stub