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An alvar is a biological environment based on a
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms w ...
plain In geography, a plain is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and as plateaus or uplands ...
with thin or no soil and, as a result, sparse grassland vegetation. Often flooded in the spring, and affected by drought in midsummer, alvars support a distinctive group of prairie-like plants. Most alvars occur either in northern Europe or around the Great Lakes in North America. This stressed habitat supports a community of rare plants and animals, including species more commonly found on
prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s.
Lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta ('' sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and ...
es are common species. Trees and bushes are absent or severely stunted. The primary cause of alvars is the shallow exposed bedrock. Flooding and drought, as noted, add to the stress of the site and prevent many species from growing. Disturbance may also play a role. In Europe, grazing is frequent, while in North America, there is some evidence that fire may also prevent encroachment by forest. The habitat also has strong competition gradients, with better competitors occupying the deeper soil and excluding other species to less productive locations. Crevices in the limestone provide a distinctive habitat which is somewhat protected from grazing, and which may provide habitat for unusual ferns such as '' Pellaea atropurpurea''. Bare rock flats provide areas with extremely low competition that serve as refugia for weak competitors such as the
sandwort Sandwort is the common name of several flowering plants in the carnation family, which may be members of the following genera: *'' Arenaria'' *''Minuartia'' *''Moehringia ''Moehringia'' is a genus of flowering plants in the family Caryophyllacea ...
'' Minuartia michauxii'' and '' Micranthes virginiensis''. In a representative set of four Ontario alvars, seven habitat types were described. From deep to shallow soil these were: tall grassy meadows, tall
forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
-rich meadows, low grassy meadows, low forb-rich meadows, dry grassland, rock margin grassland and bare rock flats. Alvars comprise a small percentage of the Earth's
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s by land extent. Although some 120 exist in the Great Lakes region, in total there are only about left across the entire Great Lakes basin, and many of these have been degraded by agriculture and other human uses. More than half of all remaining alvars occur in Ontario. There are smaller areas in New York, Michigan, Ohio, Wisconsin and Quebec. In
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, alvars provide habitat for birds such as
bobolink The bobolink (''Dolichonyx oryzivorus'') is a small New World blackbird and the only member of the genus ''Dolichonyx''. An old name for this species is the "rice bird", from its tendency to feed on cultivated grains during winter and migration. ...
s,
eastern meadowlark The eastern meadowlark (''Sturnella magna'') is a medium-sized blackbird (family: Icteridae), very similar in appearance to sister species western meadowlark. It occurs from eastern North America to northern South America, where it is also most w ...
s, upland sandpipers, eastern towhees,
brown thrasher The brown thrasher (''Toxostoma rufum''), sometimes erroneously called the brown thrush or fox-coloured thrush, is a bird in the family Mimidae, which also includes the New World catbirds and mockingbirds. The brown thrasher is abundant through ...
s and
loggerhead shrike The loggerhead shrike (''Lanius ludovicianus'') is a passerine bird in the family Laniidae. It is the only member of the shrike family endemic to North America; the related northern shrike (''L. borealis'') occurs north of its range, however it ...
s whose habitat is declining elsewhere. Rare plants include Kalm's lobelia (''Lobelia kalmii''), Pringle's aster (''Symphyotrichum pilosum'' var. ''pringlei''), juniper sedge ('' Carex juniperorum''), lakeside daisy (''
Hymenoxys acaulis ''Tetraneuris acaulis'' is a North American species of flowering plants in the sunflower family.Tetraneuris acaulis'. ITIS.Tetraneuris acaulisLady Bird Johnson Wildflower Center'', University of Texas.Hymenoxys acaulis'. '' The Jepson Manual, Uni ...
''), ram's-head lady's-slipper (''Cypripedium arietinum''), and dwarf lake iris (''Iris lacustris''). Also associated with alvars are rare
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
and
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class ...
s. The use of the word "alvar" to refer to this type of environment originated in
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
. The largest alvar in Europe is located on the Swedish island of
Öland Öland (, ; ; sometimes written ''Øland'' in other Scandinavian languages, and often ''Oland'' internationally; la, Oelandia) is the second-largest Swedish island and the smallest of the traditional provinces of Sweden. Öland has an area ...
. Here the thin soil mantle is only 0.5 to 2.0 centimeters thick in most places and in many extents consists of exposed limestone slabs. The landscape there has been designated a
UNESCO The United Nations Educational, Scientific and Cultural Organization is a List of specialized agencies of the United Nations, specialized agency of the United Nations (UN) aimed at promoting world peace and security through international coope ...
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for ...
. There are other more local names for similar landforms, such as a pavement barren, although this term is also used for similar
landform A landform is a natural or anthropogenic land feature on the solid surface of the Earth or other planetary body. Landforms together make up a given terrain, and their arrangement in the landscape is known as topography. Landforms include hills, ...
s based on
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicat ...
. In the United Kingdom the exposed landform is called a
limestone pavement A limestone pavement is a natural karst landform consisting of a flat, incised surface of exposed limestone that resembles an artificial pavement. The term is mainly used in the UK and Ireland, where many of these landforms have developed dis ...
and thinly covered limestone is known as
calcareous grassland Calcareous grassland (or alkaline grassland) is an ecosystem associated with thin basic soil, such as that on chalk and limestone downland. Plants on calcareous grassland are typically short and hardy, and include grasses and herbs such as clover. ...
.


European alvar locations

*
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
**
Öland Öland (, ; ; sometimes written ''Øland'' in other Scandinavian languages, and often ''Oland'' internationally; la, Oelandia) is the second-largest Swedish island and the smallest of the traditional provinces of Sweden. Öland has an area ...
 –
Stora Alvaret Stora Alvaret (; "the Great Alvar") is an alvar, a barren limestone terrace, in the southern half of the island of Öland, Sweden. Stora Alvaret is a dagger shaped expanse almost long and about at the widest north end. The area of this formation ...
 – largest alvar extent in Europe **
Gotland Gotland (, ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a province, county, municipality, and diocese. The province includes the islands of Fårö and Gotska Sandön to ...
**
Västergötland Västergötland (), also known as West Gothland or the Latinized version Westrogothia in older literature, is one of the 25 traditional non-administrative provinces of Sweden (''landskap'' in Swedish), situated in the southwest of Sweden. Väs ...
 – several locations on limestone mountain
Kinnekulle Kinnekulle is a flat-topped mountain in the county of Västergötland, Sweden, on the eastern shore of lake Vänern. Its highest point is above sea level. The mountain is long and wide at the top. Geology Despite its enormous size, Kinnekul ...
, smaller fragments on
Falbygden Falbygden is a geographical area, centered at the town of Falköping in Västergötland, in southwestern Sweden, and covered mostly by farmland. Most of the area belongs to Falköping Municipality and the west part of Tidaholm Municipality. In me ...
, e.g. in Dala and Högstena parishes *
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, an ...
** Alvars are distributed along the whole northern coast of Estonia from approx. the town of Paldiski to Sillamäe, wherever limestone comes to the surface near the seashore (see
Baltic Klint The Baltic Klint (Clint, Glint; et, Balti klint, sv, Baltiska klinten, russian: Балтийско-Ладожский уступ, Глинт) is an erosional limestone escarpment and cuesta on several islands of the Baltic Sea, in Estonia, in ...
), as well as on the islands of the
West Estonian archipelago West Estonian archipelago ( et, Lääne-Eesti saarestik, also Moonsund archipelago) is a group of Estonian islands located in the Baltic Sea around Väinameri. The total area is about . The archipelago is composed of the islands Saaremaa, Hiiu ...
. Estonia used to be home to approximately one third of the world's alvars; however, the total area of alvars has decreased from 43,000
hectare The hectare (; SI symbol: ha) is a non-SI metric unit of area equal to a square with 100- metre sides (1 hm2), or 10,000 m2, and is primarily used in the measurement of land. There are 100 hectares in one square kilometre. An acre i ...
s in the 1930s to 12,000 hectares in 2000, and approximately 9,000 hectares in 2010. Estonian alvars are home to 267
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
s, approximately one fifth of which are protected. There are also 142 species of
bryophyte The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in s ...
s and 263 species of
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship. The
Estonian government , image = , date = 24 February 1918 , state = Republic of Estonia , address = Rahukohtu 3, 15161 Tallinn, Estonia , appointed = President of Estonia , leader_title = Prime Minister of Estonia , ...
has committed itself to protect at least 9,800 hectares of the country's alvars as part of the
Natura 2000 Natura 2000 is a network of nature protection areas in the territory of the European Union. It is made up of Special Areas of Conservation and Special Protection Areas designated under the Habitats Directive and the Birds Directive, respecti ...
network. The
Loopealse Loopealse (Estonian for ''"Alvar"'', literally ''"Above Alvar"'') is a subdistrict ( et, asum) in the district of Lasnamäe, Tallinn, the capital of Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in N ...
subdistrict of
Tallinn Tallinn () is the most populous and capital city of Estonia. Situated on a bay in north Estonia, on the shore of the Gulf of Finland of the Baltic Sea, Tallinn has a population of 437,811 (as of 2022) and administratively lies in the Harju '' ...
is named after alvar. **
Vardi Nature Reserve Vardi Nature Reserve is a nature reserve in Rapla County in central Estonia. Vardi nature reserve encompasses one of the few remaining alvar forests in Estonia, an unusual landscape type today as cut-down forests normally never recover. The soi ...
in
Rapla County Rapla County ( et, Rapla maakond or ''Raplamaa'') is one of the fifteen counties of Estonia. It is situated in the western part of the country and borders Järva County to the east, Pärnu County to the south, Lääne County to the west, and Har ...
is an Estonian nature reserve especially designated to protect one of the more representative alvar areas of Estonia. *
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...
**
Cumbria Cumbria ( ) is a ceremonial and non-metropolitan county in North West England, bordering Scotland. The county and Cumbria County Council, its local government, came into existence in 1974 after the passage of the Local Government Act 1972. ...
and North
Yorkshire Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a Historic counties of England, historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other Eng ...
– under protection in the UK Biodiversity Action Plan *
Ireland Ireland ( ; ga, Éire ; Ulster-Scots: ) is an island in the North Atlantic Ocean, in north-western Europe. It is separated from Great Britain to its east by the North Channel, the Irish Sea, and St George's Channel. Ireland is the s ...
**
The Burren The Burren (; ) is a karst/ glaciokarst landscape centred in County Clare, on the west coast of Ireland.
Burr ...
, a large alvar in northwest
County Clare County Clare ( ga, Contae an Chláir) is a county in Ireland, in the Southern Region and the province of Munster, bordered on the west by the Atlantic Ocean. Clare County Council is the local authority. The county had a population of 118,81 ...
Sarve loopealne Hiiumaal.jpg, Sarve alvar in
Hiiumaa Hiiumaa (, ) is the second largest island in Estonia and is part of the West Estonian archipelago, in the Baltic Sea. It has an area of 989 km2 and is 22 km from the Estonian mainland. Its largest town is Kärdla. It is located within ...
island Väike-Pakri loopealsel.jpg, Alvar in Väike-Pakri island Österplana hed vegetation.jpg, Sparse vegetation,
Kinnekulle Kinnekulle is a flat-topped mountain in the county of Västergötland, Sweden, on the eastern shore of lake Vänern. Its highest point is above sea level. The mountain is long and wide at the top. Geology Despite its enormous size, Kinnekul ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
Limestone pavement above Malham Cove.jpg, Limestone pavement at
Malham Cove Malham Cove is a large curved limestone formation north of the village of Malham, North Yorkshire, England. It was formed by a waterfall carrying meltwater from glaciers at the end of the last Ice Age more than 12,000 years ago. Today it is a ...
, UK


Some North American alvar locations

*The ''rare'' Charitable Research Reserve –
Cambridge Cambridge ( ) is a university city and the county town in Cambridgeshire, England. It is located on the River Cam approximately north of London. As of the 2021 United Kingdom census, the population of Cambridge was 145,700. Cambridge bec ...
,
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
*
Lake Erie Lake Erie ( "eerie") is the fourth largest lake by surface area of the five Great Lakes in North America and the eleventh-largest globally. It is the southernmost, shallowest, and smallest by volume of the Great Lakes and therefore also ha ...
**
Kelley's Island Kelleys Island is both a village in Erie County, Ohio, and the island which it fully occupies in Lake Erie. The British originally called it Sandusky Island. Later the United States took it over and officially designated it as Island Number 6 ...
,
Ohio Ohio () is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Of the List of states and territories of the United States, fifty U.S. states, it is the List of U.S. states and territories by area, 34th-l ...
– North Shore Alvar State Nature Preserve **
Marblehead, Ohio Marblehead is a village in Ottawa County, Ohio, United States. The population was 903 at the 2010 census. It sits at the tip of the Marblehead Peninsula, which divides Lake Erie proper from Sandusky Bay. Marblehead is part of the area that is ...
– mostly destroyed by limestone quarrying **
Pelee Island Pelee may refer to: * Île Pelée, an island off Cherbourg-en-Cotentin, France * Pelee, Ontario, an island in Lake Erie, Canada *Point Pelee National Park, a park in Ontario, Canada *Mount Pelée, a volcano in Martinique *Peleus In Greek mytho ...
,
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
– Stone Road Alvar Nature Reserve *
Lake Huron Lake Huron ( ) is one of the five Great Lakes of North America. Hydrologically, it comprises the easterly portion of Lake Michigan–Huron, having the same surface elevation as Lake Michigan, to which it is connected by the , Straits of Mack ...
**Maxton Plains Proposed Natural Area,
Drummond Island Drummond may refer to: Places Antarctica * Drummond Peak, King Edward VII Land * Drummond Glacier, Graham Land Canada * Drummond (electoral district), a Quebec federal riding * Drummond (provincial electoral district), Quebec * Drummon ...
,
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and t ...
**Belanger Bay Alvar,
Manitoulin Island Manitoulin Island is an island in Lake Huron, located within the borders of the Canadian province of Ontario, in the bioregion known as Laurentia. With an area of , it is the largest lake island in the world, large enough that it has over 100 ...
, Ontario **Quarry Bay Nature Reserve, Manitoulin Island, Ontario **Bruce Alvar Nature Reserve,
Bruce Peninsula The Bruce Peninsula is a peninsula in Ontario, Canada, that divides Georgian Bay of Lake Huron from the lake's main basin. The peninsula extends roughly northwestwards from the rest of Southwestern Ontario, pointing towards Manitoulin Island, ...
, Ontario **Baptise Harbour Nature Reserve, Bruce Peninsula, Ontario ** Misery Bay Provincial Park, Manitoulin Island, Ontario *
Lake Michigan Lake Michigan is one of the five Great Lakes of North America. It is the second-largest of the Great Lakes by volume () and the third-largest by surface area (), after Lake Superior and Lake Huron. To the east, its basin is conjoined with that o ...
**Red Banks Alvar, Red Banks, Brown County, Wisconsin *
Lake Ontario Lake Ontario is one of the five Great Lakes of North America. It is bounded on the north, west, and southwest by the Canadian province of Ontario, and on the south and east by the U.S. state of New York. The Canada–United States border sp ...
**Carden Plain Alvar, City of Kawartha Lakes, Ontario, including
Carden Alvar Provincial Park Carden Alvar Provincial Park is a Ontario Parks, provincial park located in the Kawartha Lakes in Central Ontario, Central Ontario, Canada. Alvars are globally rare ecosystems found exclusively in Northern Europe and the Great Lakes region of Nor ...
**Chaumont Barrens Preserve, New York **Three Mile Creek Barrens, New York **
Burnt Lands Alvar The Burnt Lands is an alvar between Almonte and Ottawa near Upper Huntley, Ontario, Canada. It probably obtained its name from one of the forest fires that swept the area during early European settlement. It is possible that fires assist in c ...
, Almonte, Ontario ** Balsam Lake Indian Point Provincial Park, Ontario *
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirte ...
** Quyon **Alvar d' Aylmer *
Manitoba Manitoba ( ) is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population o ...
** Interlake


See also

* * * * * * *


References


External links

{{Commons-inline, Alvar (landform), Alvar * http://www.epa.gov/ecopage/shore/alvars/ Alkaline soils Habitats Landforms Types of soil Alpine flora Gotland Geography of Gotland County