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The alula , or bastard wing, (plural ''alulae'') is a small projection on the anterior edge of the
wing A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is e ...
of modern
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s and a few
non-avian dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutiona ...
s. The word is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "winglet"; it is the diminutive of ''ala'', meaning "wing". The alula is the freely moving first digit, a bird's "thumb", and typically bears three to five small flight feathers, with the exact number depending on the
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. There also are minor covert feathers overlying the flight feathers. Like the larger flight feathers found on the wing's trailing edge, these alula feathers are asymmetrical, with the shaft running closer to anterior edge.


Function

In most situations, the alula is held flush against the wing; however, it can be manipulated. When flying at slow speeds or landing, the bird moves its alula slightly upwards and forward, which creates a small slot on the wing's leading edge. This functions in the same way as the slats on the wing of an aircraft, allowing the wing to achieve a higher than normal angle of attack – and thus
lift Lift or LIFT may refer to: Physical devices * Elevator, or lift, a device used for raising and lowering people or goods ** Paternoster lift, a type of lift using a continuous chain of cars which do not stop ** Patient lift, or Hoyer lift, mobil ...
– without resulting in a stall. The tip of the alula forms a tiny vortex, acting similar to a
vortex generator A vortex generator (VG) is an aerodynamic device, consisting of a small vane usually attached to a lifting surface (or airfoil, such as an aircraft wing) or a rotor blade of a wind turbine.
, that forces the airflow over the wing to better bind to it. During stretching of the wing down toward the ground, the alula is abducted from the wing and can be clearly viewed. In falcons, the alula is more prominent, and provides a degree of control when the wing is cupped or tucked in a dive/stoop. The Alulae are particularly notable in peregrine falcons.


In ancient birds

The presence of an alula has been confirmed in several now- extinct ancient relatives of modern birds, including ''
Eoalulavis hoyasi ''Eoalulavis'' (from the Ancient Greek: ''Éōs'', "dawn"; ''alula'', "bastard wing"; ''avis'', "bird") is a monotypic genus of enantiornithean bird that lived during the Barremian, in the Lower Cretaceous around 125 million years ago. The only ...
'' (an enantiornithine from the mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
, 115 mya) and the earlier ''
Protopteryx fengningensis ''Protopteryx'' is an extinct bird and the most basal enantiornithean, from the Cretaceous period.Zhou, Zhonghe. (2002). "A new and primitive enantiornithine bird from the Early Cretaceous of China." ''Journal of Vertebrate Paleontology'', 22(1): ...
''. Since these
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are not closely related to modern birds, either the alula
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
twice, or it did so more than 130 million years ago.


See also

* Alular digit homology *
Bird flight Bird flight is the primary mode of locomotion used by most bird species in which birds take off and fly. Flight assists birds with feeding, breeding, avoiding predators, and migrating. Bird flight is one of the most complex forms of locomo ...
*
Leading edge slot A leading-edge slot is a fixed aerodynamic feature of the wing of some aircraft to reduce the stall speed and promote good low-speed handling qualities. A leading-edge slot is a spanwise gap in each wing, allowing air to flow from below the wing ...
, the man-made equivalent on fixed-wing aircraft


Citations


References

* * * {{Birds Feathers Bird anatomy