African harrier hawk
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The African harrier-hawk, harrier hawk or gymnogene (''Polyboroides typus'') is a bird of prey. It is about in length. It breeds in most of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
south of the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
. The only other member of the genus is the
allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
Madagascar harrier-hawk (''Polyboroides radiatus'').


Description

left, 165px, Adult hunting at a weaver colony in Etosha National Park, Namibia The African harrier-hawk is a medium-sized raptor. The upperparts, head and breast are pale grey. The belly is white with fine dark barring. The broad wings are pale grey with a black trailing edge fringed with a narrow white line. The tail is black with a single broad white band. There is a bare facial patch of variable colour, usually red or yellow. Genders are similar, but young birds have pale brown instead of grey, and dark brown replacing black. An unusual trait of this species is the double-jointed knees it possesses, which enable it to reach into otherwise inaccessible holes and cracks for prey. A comparable leg-structure and behaviour can be found in the Neotropical crane hawk as well as the extinct
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
n '' Pengana''; a case of convergent evolution. The call is a whistled '.


Distribution and habitat

African harrier-hawks are a common raptorial species south of the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
being most commonly found in the tropical regions of western
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
becoming less common in East and South Africa. African harrier-hawks are adaptable in their habitat preferences, occupying the following habitats in the Dzanga-Sangha Special Reserve in the Central African Republic: thick
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...
, forest edge, riparian areas, agricultural land and human occupied areas. African harrier-hawks are adaptable and able to live in both urban and rural human occupied areas and they are one of the most common raptorial species in traditional rural villages of eastern Guinea-Bissau. African harrier-hawks have also been known to breed in Palm trees present in cities and urban gardens.


Biology


Breeding


Breeding season

The breeding season starts at different times in different parts of African harrier-hawk distribution. In Nigeria the breeding season is in March-August and South of the
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can als ...
it appears that the breeding season is in the austral summer but can vary in the months of different countries in South Africa it is November-December but in Zambia, Malawi and Zimbabwe it is September-November.


Nests and nesting

Nest sites are most commonly in large trees that are sometimes growing out of or located on a rocky outcrop. Nests are circular and often placed in the main fork of the tree and are below the canopy. Nests can be used for several breeding seasons and are relatively big as are other raptors reaching estimated sizes of 0.75m wide and 0.2m deep. Nests are made of sticks and are lined with leaves from trees neighbouring the nest. The clutch is one to three eggs.


Courtship

In the courtship display for African harrier-hawks either one of or both individuals in a pair soar slowly together, at height, and can often be heard calling during this time. When the male flies on his own he often flies in an undulating pattern and flaps his wings. When the pair fly together, there have been records of the male diving towards the female and touching her back with his
talon Talon or talons may refer to: Science and technology * Talon (anatomy), the claw of a bird of prey * Brodifacoum, a rodenticide, also known as the brand Talon * TALON (database), a database maintained by the US Air Force * Talon, an anti-vehicle- ...
s, and the female turning over and touching talons with the male.


Foraging


Diet

The African harrier-hawk is omnivorous, eating the fruit of the oil palm as well as hunting small vertebrates. Its ability to climb, using wings as well as feet, and its long double-jointed legs, enable this bird to raid the nests of cavity-nesters such as
barbets Barbet may refer to: * Barbet (dog), a dog breed * Various birds in the infraorder Ramphastides ** African barbet, part of the bird family Lybiidae ** New World barbet, the bird family Capitonidae ** Asian barbet The Asian barbets are a family ...
and woodhoopoes for eggs and nestlings. It has been known to prey on introduced species such as feral pigeons,
house sparrow The house sparrow (''Passer domesticus'') is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, a ...
s and eastern gray squirrels.


Foraging techniques

African harrier-hawks have been identified to employ four different hunting strategies namely: low soaring, high soaring, perch hunting and, canopy and ground foraging. Low soaring is the most commonly used method The harrier-hawk flies close to the canopy and is often mobbed by small passerine birds. The African harrier-hawk uses the level of aggression shown to help locate nest sites of these passerines and has been observed to turn around when the mobbing by a passerine becomes less aggressive, the harrier-hawk will begin looking for nest sites in the trees once it has found the area where the passerines show the most aggression towards the African harrier-hawk. To catch reptiles in the open African harrier-hawks use high soaring, flying at a maximum of 100m. They descend quickly to a height just above that of the vegetation to where the prey was located. Perch hunting is often used to hunt invertebrates such as
orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...
and other insects. Canopy and ground foraging is where the harrier-hawk either walks on the ground or moves between branches in the canopy looking for prey, looking into crevices and holes in both trees and on the ground.


Gallery

African Harrier-hawk (Polyboroides typus) (21646641084).jpg, Immature bird in flight, Zambia Immature African harrier-hawk (gymnogene), Polyboroides typus, at Ndumo Nature Reserve, KwaZulu-Natal, South Africa (28901676521).jpg, Immature bird lifting tree bark, South Africa African harrier-hawk (Polyboroides typus typus) taking off, crop.jpg, Adult taking flight, Zimbabwe Polyboroides typus -Lake Nakuru National Park, Kenya-8 (2).jpg, Adult investigating a tree cavity, Kenya Martial eagle (Polemaetus bellicosus) in flight with African harrier-hawk (Polyboroides typus) composite.jpg, Adult soaring alongside a martial eagle (''
Polemaetus bellicosus The martial eagle (''Polemaetus bellicosus'') is a large eagle native to sub-Saharan Africa.Ferguson-Lees & Christie, ''Raptors of the World''. Houghton Mifflin Company (2001), . It is the only member of the genus ''Polemaetus''. A species of ...
)'', Zimbabwe


References


External links

* (African harrier-hawk = ) Gymnogene
Species text in The Atlas of Southern African Birds
{{DEFAULTSORT:harrier-hawk, African African harrier-hawk African harrier-hawk African harrier-hawk Taxa named by Andrew Smith (zoologist)