African harrier-hawk
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The African harrier-hawk, harrier hawk or gymnogene (''Polyboroides typus'') is a
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
. It is about in length. It breeds in most of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
south of the Sahara. The only other member of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
is the allopatric
Madagascar harrier-hawk The Madagascar harrier-hawk (''Polyboroides radiatus'') is a very large species of bird of prey in the family Accipitridae, endemic to Madagascar. Description The Madagascar harrier-hawk is a largish raptor with long and broad wings which when f ...
(''Polyboroides radiatus'').


Description

left, 165px, Adult hunting at a weaver colony in Etosha National Park, Namibia The African harrier-hawk is a medium-sized raptor. The upperparts, head and breast are pale grey. The belly is white with fine dark barring. The broad wings are pale grey with a black trailing edge fringed with a narrow white line. The tail is black with a single broad white band. There is a bare facial patch of variable colour, usually red or yellow. Genders are similar, but young birds have pale brown instead of grey, and dark brown replacing black. An unusual trait of this species is the double-jointed knees it possesses, which enable it to reach into otherwise inaccessible holes and cracks for prey. A comparable leg-structure and behaviour can be found in the
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
crane hawk The crane hawk (''Geranospiza caerulescens'') is a species of bird of prey in the family Accipitridae. It is monotypic within the genus ''Geranospiza''. Taxonomy The crane hawk used to be many species that were recently lumped into one. Those ...
as well as the extinct Australian ''
Pengana ''Pengana robertbolesi'', also referred to as the flexiraptor, is an extinct bird of prey that lived during the Early Miocene (23–16 million years ago). Living relatives of ''P. robertbolesi'' may include the harriers. When alive, it may ...
''; a case of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. The call is a whistled '.


Distribution and habitat

African harrier-hawks are a common raptorial species south of the Sahara being most commonly found in the tropical regions of western
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
becoming less common in East and
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. African harrier-hawks are adaptable in their habitat preferences, occupying the following habitats in the Dzanga-Sangha Special Reserve in the
Central African Republic The Central African Republic (CAR; ; , RCA; , or , ) is a landlocked country in Central Africa. It is bordered by Chad to the north, Sudan to the northeast, South Sudan to the southeast, the DR Congo to the south, the Republic of th ...
: thick rainforest, forest edge,
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
areas, agricultural land and human occupied areas. African harrier-hawks are adaptable and able to live in both urban and rural human occupied areas and they are one of the most common raptorial species in traditional
rural In general, a rural area or a countryside is a geographic area that is located outside towns and cities. Typical rural areas have a low population density and small settlements. Agricultural areas and areas with forestry typically are descri ...
villages of eastern
Guinea-Bissau Guinea-Bissau ( ; pt, Guiné-Bissau; ff, italic=no, 𞤘𞤭𞤲𞤫 𞤄𞤭𞤧𞤢𞥄𞤱𞤮, Gine-Bisaawo, script=Adlm; Mandinka: ''Gine-Bisawo''), officially the Republic of Guinea-Bissau ( pt, República da Guiné-Bissau, links=no ), ...
. African harrier-hawks have also been known to breed in Palm trees present in cities and
urban Urban means "related to a city". In that sense, the term may refer to: * Urban area, geographical area distinct from rural areas * Urban culture, the culture of towns and cities Urban may also refer to: General * Urban (name), a list of people ...
gardens.


Biology


Breeding


Breeding season

The breeding season starts at different times in different parts of African harrier-hawk distribution. In
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
the breeding season is in March-August and South of the equator it appears that the breeding season is in the
austral summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, ...
but can vary in the months of different countries in South Africa it is November-December but in
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are t ...
,
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeas ...
and
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and ...
it is September-November.


Nests and nesting

Nest sites are most commonly in large trees that are sometimes growing out of or located on a rocky outcrop. Nests are circular and often placed in the main fork of the tree and are below the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
. Nests can be used for several breeding seasons and are relatively big as are other raptors reaching estimated sizes of 0.75m wide and 0.2m deep. Nests are made of sticks and are lined with leaves from trees neighbouring the nest. The clutch is one to three eggs.


Courtship

In the courtship display for African harrier-hawks either one of or both individuals in a pair soar slowly together, at height, and can often be heard calling during this time. When the male flies on his own he often flies in an undulating pattern and flaps his wings. When the pair fly together, there have been records of the male diving towards the female and touching her back with his talons, and the female turning over and touching talons with the male.


Foraging


Diet

The African harrier-hawk is omnivorous, eating the fruit of the
oil palm ''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm '' Elaeis guineensis'' (the species name ''guineensis'' referring to its c ...
as well as hunting small vertebrates. Its ability to climb, using wings as well as feet, and its long double-jointed legs, enable this bird to raid the nests of cavity-nesters such as barbets and
woodhoopoe The wood hoopoes or scimitarbills are a small African family, Phoeniculidae, of near passerine birds. They live south of the Sahara Desert and are not migratory. While the family is now restricted to Sub-Saharan Africa, fossil evidence shows th ...
s for eggs and nestlings. It has been known to prey on introduced species such as
feral pigeon Feral pigeons (''Columba livia domestica'' or ''Columba livia forma urbana''), also called city doves, city pigeons, or street pigeons,Nagy, Kelsi, and Johnson, Phillip David. ''Trash animals: how we live with natures filthy, feral, invasive, an ...
s, house sparrows and
eastern gray squirrel The eastern gray squirrel (''Sciurus carolinensis''), also known, particularly outside of North America, as simply the grey squirrel, is a tree squirrel in the genus ''Sciurus''. It is native to eastern North America, where it is the most prodi ...
s.


Foraging techniques

African harrier-hawks have been identified to employ four different hunting strategies namely: low soaring, high soaring, perch hunting and,
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
and ground foraging. Low soaring is the most commonly used method The harrier-hawk flies close to the canopy and is often mobbed by small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
birds. The African harrier-hawk uses the level of aggression shown to help locate nest sites of these passerines and has been observed to turn around when the mobbing by a passerine becomes less aggressive, the harrier-hawk will begin looking for nest sites in the trees once it has found the area where the passerines show the most aggression towards the African harrier-hawk. To catch reptiles in the open African harrier-hawks use high soaring, flying at a maximum of 100m. They descend quickly to a height just above that of the vegetation to where the prey was located. Perch hunting is often used to hunt
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s such as orthoptera and other insects. Canopy and ground foraging is where the harrier-hawk either walks on the ground or moves between branches in the canopy looking for prey, looking into crevices and holes in both trees and on the ground.


Gallery

African Harrier-hawk (Polyboroides typus) (21646641084).jpg, Immature bird in flight, Zambia Immature African harrier-hawk (gymnogene), Polyboroides typus, at Ndumo Nature Reserve, KwaZulu-Natal, South Africa (28901676521).jpg, Immature bird lifting tree bark, South Africa African harrier-hawk (Polyboroides typus typus) taking off, crop.jpg, Adult taking flight, Zimbabwe Polyboroides typus -Lake Nakuru National Park, Kenya-8 (2).jpg, Adult investigating a tree cavity, Kenya Martial eagle (Polemaetus bellicosus) in flight with African harrier-hawk (Polyboroides typus) composite.jpg, Adult soaring alongside a martial eagle ('' Polemaetus bellicosus)'', Zimbabwe


References


External links

* (African harrier-hawk = ) Gymnogene
Species text in The Atlas of Southern African Birds
{{DEFAULTSORT:harrier-hawk, African
African harrier-hawk The African harrier-hawk, harrier hawk or gymnogene (''Polyboroides typus'') is a bird of prey. It is about in length. It breeds in most of Africa south of the Sahara. The only other member of the genus is the allopatric Madagascar harrier-haw ...
African harrier-hawk The African harrier-hawk, harrier hawk or gymnogene (''Polyboroides typus'') is a bird of prey. It is about in length. It breeds in most of Africa south of the Sahara. The only other member of the genus is the allopatric Madagascar harrier-haw ...
African harrier-hawk The African harrier-hawk, harrier hawk or gymnogene (''Polyboroides typus'') is a bird of prey. It is about in length. It breeds in most of Africa south of the Sahara. The only other member of the genus is the allopatric Madagascar harrier-haw ...
Taxa named by Andrew Smith (zoologist)