Andreas Franz Wilhelm Schimper (12 May 1856 – 9 September 1901) was a German botanist and phytogeographer who made major contributions in the fields of

histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vi ...

ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overl ...

and plant geography. He travelled to South East Asia and the Caribbean as part of the 1899 deep-sea expedition. He coined the terms

tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equa ...

and

sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...

and is commemorated in numerous specific names.

Biography

Schimper was born in

Strassburg Strasbourg (, , ; german: Straßburg ; gsw, label= Bas Rhin Alsatian, Strossburi , gsw, label= Haut Rhin Alsatian, Strossburig ) is the prefecture and largest city of the Grand Est region of eastern France and the official seat of the E ...

, (present day Strasbourg, France), into a family of eminent scientists. His father

Wilhelm Philippe Schimper Wilhelm Philippe Schimper (January 12, 1808 – March 20, 1880, in Lichtenberg) was an Alsatian botanist with French, later German citizenship. He was born in Dossenheim-sur-Zinsel, but spent his youth in Offwiller, a village at the foot of t ...

(1808-1880) was Director of the Natural History Museum in the same town, Professor of Geology, and a leading bryologist. His father's cousin was Georg Wilhelm Schimper (1804-1878), prominent collector and explorer in

Arabia The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plat ...

and

North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...

; the naturalist Karl Friedrich Schimper was also a relative. Schimper studied at the

University of Strassburg The University of Strasbourg (french: Université de Strasbourg, Unistra) is a public research university located in Strasbourg, Alsace, France, with over 52,000 students and 3,300 researchers. The French university traces its history to the ea ...

from 1874 to 1878, acquiring a Ph.D. He then worked in

Lyon Lyon,, ; Occitan language, Occitan: ''Lion'', hist. ''Lionés'' also spelled in English as Lyons, is the List of communes in France with over 20,000 inhabitants, third-largest city and Urban area (France), second-largest metropolitan area of F ...

, and in 1880 travelled to the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...

, becoming a Fellow at

Johns Hopkins University Johns Hopkins University (Johns Hopkins, Hopkins, or JHU) is a private research university in Baltimore, Maryland. Founded in 1876, Johns Hopkins is the oldest research university in the United States and in the western hemisphere. It consi ...

. In 1882, he moved back to the

University of Bonn The Rhenish Friedrich Wilhelm University of Bonn (german: Rheinische Friedrich-Wilhelms-Universität Bonn) is a public research university located in Bonn, North Rhine-Westphalia, Germany. It was founded in its present form as the ( en, Rhine ...

working with Eduard Strasburger, becoming a private docent. In 1883, Schimper postulated the

endosymbiotic An ''endosymbiont'' or ''endobiont'' is any organism that lives within the body or cells of another organism most often, though not always, in a mutualistic relationship. (The term endosymbiosis is from the Greek: ἔνδον ''endon'' "within ...

origin of

chloroplasts A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in ...

and paved the way to the

symbiogenesis Symbiogenesis (endosymbiotic theory, or serial endosymbiotic theory,) is the leading evolutionary theory of the origin of eukaryotic cells from prokaryotic organisms. The theory holds that mitochondria, plastids such as chloroplasts, and pos ...

theory of

Konstantin Mereschkowski Konstantin Sergeevich Mereschkowski ( rus, Константи́н Серге́евич Мережко́вский, p=mʲɪrʲɪˈʂkofskʲɪj; – 9 January 1921) was a prominent Russian biologist and botanist, active mainly around Kazan, Tat ...

and

Lynn Margulis Lynn Margulis (born Lynn Petra Alexander; March 5, 1938 – November 22, 2011) was an American evolutionary biologist, and was the primary modern proponent for the significance of symbiosis in evolution. Historian Jan Sapp has said that "Lynn Ma ...

. In 1886, he was appointed Extraordinary Professor at the University of Bonn, and worked largely on cell histology,

chromatophores Chromatophores are cells that produce color, of which many types are Biological pigment, pigment-containing cells, or groups of cells, found in a wide range of animals including amphibians, fish, reptiles, crustaceans and cephalopods. Mammals and ...

and starch metabolism. He had become interested in

phytogeography Phytogeography (from Greek φυτόν, ''phytón'' = "plant" and γεωγραφία, ''geographía'' = "geography" meaning also distribution) or botanical geography is the branch of biogeography that is concerned with the geographic distribution ...

and plant ecology, undertaking expeditions to the

West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greate ...

and

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

in 1882–1883. In 1886, he stayed with

Fritz Müller Johann Friedrich Theodor Müller (31 March 1822 – 21 May 1897), better known as Fritz Müller, and also as Müller-Desterro, was a German biologist who emigrated to southern Brazil, where he lived in and near the German community of Blumenau, ...

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

, and in 1889–1890 in Ceylon, the Malaya and Botanical Garden in

Buitenzorg Bogor ( su, , nl, Buitenzorg) is a city in the West Java province, Indonesia. Located around south of the national capital of Jakarta, Bogor is the 6th largest city in the Jakarta metropolitan area and the 14th overall nationwide.Bogor,

Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mos ...

), concentrating on mangroves, epiphytes and

littoral The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal a ...

vegetation. This resulted in his account of the

Rhizophoraceae The Rhizophoraceae is a family of tropical or subtropical flowering plants. It includes around 147 species distributed in 15 genera.Setoguchi, H., Kosuge, K., & Tobe, H. (1999). Molecular Phylogeny of Rhizophoraceae Based on rbcL Gene Sequences. ...

in Engler & Prantl's ''Natürliche Pflanzenfamilien''. In 1898, he became Professor of Botany at the

University of Basel The University of Basel (Latin: ''Universitas Basiliensis'', German: ''Universität Basel'') is a university in Basel, Switzerland. Founded on 4 April 1460, it is Switzerland's oldest university and among the world's oldest surviving universit ...

and the same year joined the

German German(s) may refer to: * Germany (of or related to) ** Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...

Valdivia-Expedition. This was a deep-sea expedition aboard the SS ''Valdivia'' led by Professor Carl Chun. The trip lasted 9 months, during which they visited the Canary Islands,

Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...

Cape Town Cape Town ( af, Kaapstad; , xh, iKapa) is one of South Africa's three capital cities, serving as the seat of the Parliament of South Africa. It is the legislative capital of the country, the oldest city in the country, and the second largest ...

, (where he joined

Rudolf Marloth Hermann Wilhelm Rudolf Marloth (28 December 1855 Lübben, Germany – 15 May 1931 Caledon, Cape Province) was a German-born South African botanist, pharmacist and analytical chemist, best known for his ''Flora of South Africa'' which appeared i ...

on collecting trips in the southern Cape),

Kerguelen The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...

, New Amsterdam and

Cocos Islands ) , anthem = "''Advance Australia Fair''" , song_type = , song = , image_map = Australia on the globe (Cocos (Keeling) Islands special) (Southeast Asia centered).svg , map_alt = Location of the Cocos (Keeling) Islands , map_caption = ...

, Sumatra, the

Maldives Maldives (, ; dv, ދިވެހިރާއްޖެ, translit=Dhivehi Raajje, ), officially the Republic of Maldives ( dv, ދިވެހިރާއްޖޭގެ ޖުމްހޫރިއްޔާ, translit=Dhivehi Raajjeyge Jumhooriyyaa, label=none, ), is an archipelag ...

, Ceylon, the

Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...

and the

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...

. Schimper was the first to give a clear and proper definition of plastid and also explained its types. In 1899, he became Professor of Botany at the

. His health had been seriously affected by

malaria Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. S ...

contracted in

and

Dar-es-Salaam Dar es Salaam (; from ar, دَار السَّلَام, Dâr es-Selâm, lit=Abode of Peace) or commonly known as Dar, is the largest city and financial hub of Tanzania. It is also the capital of Dar es Salaam Region. With a population of over s ...

, and he died of complications of malaria at the age of 45 in 1901.

Publications

Schimper is best known for ''Pflanzengeographie auf physiologischer Grundlage'', published at the

University of Jena The University of Jena, officially the Friedrich Schiller University Jena (german: Friedrich-Schiller-Universität Jena, abbreviated FSU, shortened form ''Uni Jena''), is a public research university located in Jena, Thuringia, Germany. The un ...

in 1898 where he aimed to explain the expansion and ecology of plants based on the ecological knowledge of the time. In this book he coined the terms

and

. He wrote in the preface: "''Nur wenn sie in engster Fühlung mit der experimentellen Physiologie verbleibt, wird die Ökologie der Pflanzengeographie neue Bahnen eröffnen können, denn sie setzt eine genaue Kenntnis der Lebensbedingungen der Pflanze voraus, welche nur das Experiment verschaffen kann''"''Plant-geography upon a physiological basis''
A.F.W. Schimper, 1903, Biodiversity Heritage Library(1898: IV). His classification of plant formations was important for the development of the botanical sciences: „''Nach dem Vorhergehenden sind zwei ökologische Formationsgruppen zu unterscheiden, die klimatischen oder Gebietsformationen, deren Vegetationscharakter durch die Hydrometeore beherrscht, und die edaphischen oder Standortsformationen, wo derselbe in erster Linie durch die Bodenbeschaffenheit bedingt ist''“ (1898:175–176). At the same time as his Russian

soil science Soil science is the study of soil as a natural resource on the surface of the Earth including soil formation, classification and mapping; physical, chemical, biological, and fertility properties of soils; and these properties in relation to th ...

colleagues, Schimper discussed the hypothesis of vegetation being limited to climate zones versus those that are

azonal In geography, azonal is an adjective that refers to processes or things that are not restricted to any climate zone. It can be used to describe soils, landforms, geomorphic processes or vegetation. Volcanism and mountain-related processes are exampl ...

, which was later elaborated by Frederic Edward Clements (1916) and geobotanist

Heinrich Walter Heinrich Karl Walter (21 October 1898 – 15 October 1989) was a German-Russian botanist and eco-physiologist. Life Walter, the son of a doctor, was born in Odessa. He studied plant biology at the University of Odessa from 1915 to 1917. In 1918 ...

(1954) amongst others. In 1894, Schimper was one of the 4 original authors of the textbook of botany ''Lehrbuch der Botanik'' (named. "Strasburger") and until the 5th edition 1902 editor of the chapter spermatophyta or seed-bearing plants.

wrote an account of the Cape floral region for Chun's proposed ''Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer Valdivia 1898-1899.'' and Schimper contributed two chapters on "Gebiet der Hartlaubgehölze" (Region of hardwood trees) and "Der Knysnawald".

Honours and distinctions

Schimper is commemorated in specific names such as '' Acokanthera schimperi'' and '' Harpachne schimperi''. In 1892, he was voted a member of the

Deutsche Akademie der Naturforscher Leopoldina The German National Academy of Sciences Leopoldina (german: Deutsche Akademie der Naturforscher Leopoldina – Nationale Akademie der Wissenschaften), short Leopoldina, is the national academy of Germany, and is located in Halle (Saale). Founded ...

gewählt.Mitgliederverzeichnis Leopoldina, Wilhelm Schimper
/ref>

References

External links

* *
Books by and about A.F.G. Schimper on WorldCat

Digital edition: "Anleitung zur mikroskopischen Untersuchung der vegetabilischen Nahrungs- und Genussmittel" 2nd ed. (1900)
by the University and State Library Düsseldorf
''Plant-geography upon a physiological basis''
by A.F.W. Schimper, 1903 Biodiversity Heritage Library *''Botanical Exploration of Southern Africa'' Mary Gunn & LE Codd (Balkema 1981) {{DEFAULTSORT:Schimper, Andreas Franz Wilhelm 1856 births 1901 deaths Alsatian-German people 19th-century German botanists Botanists active in the Caribbean Botanists active in South America German ecologists French histologists German histologists German phytogeographers Scientists from Strasbourg Symbiogenesis researchers University of Strasbourg alumni University of Bonn faculty University of Basel faculty Members of the German Academy of Sciences Leopoldina