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Alouatta
Howler monkeys (genus ''Alouatta'', monotypic in subfamily Alouattinae) are the most widespread primate genus in the Neotropics and are among the largest of the platyrrhines along with the muriquis (''Brachyteles''), the spider monkeys (''Ateles'') and woolly monkeys (''Lagotrix''). These monkeys are native to South and Central American forests. They are famous for their loud howls, which can travel more than a mile through dense rain forest. Fifteen species are recognized. Previously classified in the family Cebidae, they are now placed in the family Atelidae. They are primarily folivores but also significant frugivores, acting as seed dispersal agents through their digestive system and their locomotion. Threats include human predation, habitat destruction, and capture for pets or zoo animals. Classification * ''A. palliata'' group ** Coiba Island howler, ''Alouatta coibensis'' *** ''Alouatta coibensis coibensis'' *** Azuero howler, ''Alouatta coibensis trabeata'' ** Mantl ...
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Mantled Howler
The mantled howler (''Alouatta palliata'') is a species of howler monkey, a type of New World monkey, from Central and South America. It is one of the monkey species most often seen and heard in the wild in Central America. It takes its "mantled" name from the long guard hairs on its sides. The mantled howler is one of the largest Central American monkeys, and males can weigh up to . It is the only Central American monkey that eats large quantities of leaves; it has several adaptations to this folivorous diet. Since leaves are difficult to digest and provide less energy than most foods, the mantled howler spends the majority of each day resting and sleeping. The male mantled howler has an enlarged hyoid bone, a hollow bone near the vocal cords, which amplifies the calls made by the male, and is the reason for the name "howler". Howling allows the monkeys to locate each other without expending energy on moving or risking physical confrontation. The mantled howler lives in gro ...
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Atelidae
The Atelidae are one of the five family (biology), families of New World monkeys now recognised. It was formerly included in the family Cebidae. Atelids are generally larger monkeys; the family includes the howler monkey, howler, spider monkey, spider, woolly monkey, woolly, and Muriqui, woolly spider monkeys (the latter being the largest of the New World monkeys). They are found throughout the forested regions of Central America, Central and South America, from Mexico to northern Argentina. Characteristics The Atelidae family consists of monkeys that are small to moderate in size, usually 34 to 72 cm in head-body length, with the howler monkeys being the largest members of the group, and the spider monkeys being the smallest. They have long prehensile tails with a sensitive, almost hairless, tactile pad on the underside of the Anatomical terms of location#Proximal and distal, distal part. The tail is frequently used as 'fifth limb' while moving through the trees where they ...
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Mexican Howler Monkey
The Mexican howler (''Alouatta palliata mexicana'') is a subspecies of the mantled howler, ''A. palliata''. This subspecies is found predominantly in forests between south eastern Mexico and north eastern Peru. Typical of its species, the Mexican howler monkey has a prehensile tail, a deep jaw, and a large pharynx which it uses to make characteristically deep and resonating howls. Mantled howler monkeys are known for forming unusually large cohorts averaging 14 members and sometimes extending to 40 members. Taxonomy There are five subspecies of the ''A. palliata'' which are ''Alouatta palliata aequatorialis'', ''Alouatta palliata coibensis'', ''Alouatta palliata palliata'', ''Alouatta palliata trabeata'' and ''Alouatta palliata mexicana''. Compared with members of the howler monkey genus, the Mexican howler is sympatric with the Guatemalan black howler, ''A. pigra'', in Tabasco, Mexico. The Mexican howler differs from the golden-mantled howler, ''A. palliata palliata'', primaril ...
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Red-handed Howler
The red-handed howler (''Alouatta belzebul'') is a vulnerable species of howler monkey, a type of New World monkey. It is endemic to Brazil, found in the southeastern Amazon and disjunctly in the Atlantic Forest between Rio Grande do Norte and Sergipe. Taxonomy Considerable taxonomic confusion has surrounded this species. Until 2001, most authorities included the Amazon black howler as a subspecies (or simply a taxonomically insignificant variation) of the red-handed howler, Groves, C. (2001). ''Primate Taxonomy.'' Smithsonian Institution Press. though its distinction had already been pointed out much earlier.da Cruz Lima, E. (1945). ''Mammals of Amazônia.'' Vol. 1. Contribuições do Museu Paraense Emílio Goeldi de História Natural e Etnografia. The red-handed howler remained variable in ecology, colour and pattern of the fur, shape of the cranium, and shape of the hyoid bone (of great importance in the voice of the howler monkeys; a likely isolating mechanism between ...
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Red-handed Howler
The red-handed howler (''Alouatta belzebul'') is a vulnerable species of howler monkey, a type of New World monkey. It is endemic to Brazil, found in the southeastern Amazon and disjunctly in the Atlantic Forest between Rio Grande do Norte and Sergipe. Taxonomy Considerable taxonomic confusion has surrounded this species. Until 2001, most authorities included the Amazon black howler as a subspecies (or simply a taxonomically insignificant variation) of the red-handed howler, Groves, C. (2001). ''Primate Taxonomy.'' Smithsonian Institution Press. though its distinction had already been pointed out much earlier.da Cruz Lima, E. (1945). ''Mammals of Amazônia.'' Vol. 1. Contribuições do Museu Paraense Emílio Goeldi de História Natural e Etnografia. The red-handed howler remained variable in ecology, colour and pattern of the fur, shape of the cranium, and shape of the hyoid bone (of great importance in the voice of the howler monkeys; a likely isolating mechanism between ...
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Ecuadorian Mantled Howler
The Ecuadorian mantled howler (''Alouatta palliata aequatorialis'') is a subspecies of the mantled howler, ''A. palliata''. It ranges from Panama (or possibly the eastern tip of Costa Rica) through Colombia and Ecuador into northern Peru. The range limits between the Ecuadorian mantled howler and the golden-mantled howler The golden-mantled howler (''Alouatta palliata palliata'') is a subspecies of the mantled howler, ''A. palliata''. It ranges throughout much of Central America, in Guatemala, Honduras, Nicaragua and Costa Rica, and possibly Panama. The range lim ... are not entirely clear. The Ecuadorian mantled howler replaces the Golden-mantled howler in either extreme eastern Costa Rica or western Panama. The Ecuadorian mantled howler differs from the golden-mantled howler primarily by being paler, with a more yellowish mantle. File:Panamanian Male Adult Howler Monkey.jpg, male File:Ecuadorian mantled howler (Alouatta palliata aequatorialis) family group.jpg, troop Fil ...
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Alouatta Coibensis Coibensis
The Coiba Island howler (''Alouatta coibensis'') is a type of howler monkey, a type of New World monkey, endemic to Panama. Although the Coiba Island howler has been recognized as a separate species by a number of authorities since a 1987 study of its fingerprints, mitochondrial DNA testing found it does not differ from mantled howler populations in any significant way. A reason given for treating it as a separate species is that the dermal ridges of its hands and feet differ from those of the mantled howler. ''A. c. coibensis'' is smaller than other Central American howler monkeys and has duller pelage than the Azuero howler, ''Alouatta coibensis trabeata''. Subspecies Two subspecies of this howler have been recognized by those who consider it a separate species: *''Alouatta coibensis coibensis'' Thomas, 1902, found on Coiba Island and Jicaron, off the Pacific coast of Panama *Azuero howler, ''A. c. trabeata'' Lawrence, 1933, endemic to the Azuero Peninsula Azuero Peninsul ...
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Coiba Island Howler
The Coiba Island howler (''Alouatta coibensis'') is a type of howler monkey, a type of New World monkey, endemic to Panama. Although the Coiba Island howler has been recognized as a separate species by a number of authorities since a 1987 study of its fingerprints, mitochondrial DNA testing found it does not differ from mantled howler populations in any significant way. A reason given for treating it as a separate species is that the dermal ridges of its hands and feet differ from those of the mantled howler. ''A. c. coibensis'' is smaller than other Central American howler monkeys and has duller pelage than the Azuero howler, ''Alouatta coibensis trabeata''. Subspecies Two subspecies of this howler have been recognized by those who consider it a separate species: *''Alouatta coibensis coibensis'' Thomas, 1902, found on Coiba Island and Jicaron, off the Pacific coast of Panama *Azuero howler, ''A. c. trabeata'' Lawrence, 1933, endemic to the Azuero Peninsula Azuero Peninsula ( ...
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Ursine Howler
The ursine howler (''Alouatta arctoidea'') is a species of howler monkey native to Venezuela and possibly Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the .... It is sometimes considered a subspecies of the Venezuelan red howler and classified as ''Alouatta seniculus arctoidea''. References ursine howler Mammals of Venezuela ursine howler Taxa named by Ángel Cabrera (naturalist) {{newworld-monkey-stub ...
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Golden-mantled Howler
The golden-mantled howler (''Alouatta palliata palliata'') is a subspecies of the mantled howler, ''A. palliata''. It ranges throughout much of Central America, in Guatemala, Honduras, Nicaragua and Costa Rica, and possibly Panama. The range limits between the golden-mantled howler and the Ecuadorian mantled howler are not entirely clear. The Ecuadorian mantled howler replaces the golden-mantled howler in either extreme eastern Costa Rica or western Panama. The golden-mantled howler differs from the Ecuadorian mantled howler primarily by being darker, with a mantle that is more rufous than yellowish. The golden-mantled howler differs from the Mexican howler monkey primarily in aspects of skull morphology. References golden-mantled howler Primates of Central America golden-mantled howler golden-mantled howler The golden-mantled howler (''Alouatta palliata palliata'') is a subspecies of the mantled howler, ''A. palliata''. It ranges throughout much of Central America, in ...
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Azuero Howler
The Azuero howler (''Alouatta coibensis trabeata'') a type of monkey that is a subspecies of the Coiba Island howler ''A. coibensis''. This subspecies is endemic to the Azuero Peninsula in Panama. The Azuero howler is distinguished primarily by its golden flanks and loins, and browner appearance on the rest of its body. Although generally considered a subspecies of ''A. coibensis'', there is some debate within the scientific community as to whether ''A. coibensis'' itself is a valid species. If not, ''A. c. trabeata'' would be considered subspecies of the mantled howler The mantled howler (''Alouatta palliata'') is a species of howler monkey, a type of New World monkey, from Central and South America. It is one of the monkey species most often seen and heard in the wild in Central America. It takes its "mantle ..., ''A. palliata''. In that case its trinomial name would be ''A. p. trabeata''. References Primates of Central America Howler monkeys Mammals described in ...
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Howling
Howling is a vocal form of animal communication seen in most canines, particularly wolves, coyotes, foxes, and dogs, as well as cats and some species of monkeys. Howls are generally lengthy sustained sounds, loud and audible over long distances, often with some variation in pitch over the length of the sound. Howling is generally used by animals that engage in this behavior to signal their positions to one another, to call the pack to assemble, or to note their territory. The behavior is occasionally copied by humans, and has been noted to have varying degrees of significance in human culture. In canines The long-distance howling of wolves and coyotes is one way in which canines communicate. Long-distance contact calls are common in Canidae, typically in the form of either barks (termed "pulse trains") or howls (termed "long acoustic streams"). Wolves howl to assemble the pack usually before and after hunts, to pass on an alarm particularly at a den site, to locate each ...
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