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Rosids
The ROSIDS are members of a large clade (monophyletic group) of flowering plants , containing about 70,000 species , more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders , depending upon circumscription and classification . These orders, in turn, together comprise about 140 families . Fossil rosids are known from the Cretaceous
Cretaceous
period. Molecular clock estimates indicate that the rosids originated in the Aptian or Albian stages of the Cretaceous, between 125 and 99.6 million years ago. CONTENTS * 1 Name * 2 Relationships * 3 Classification * 3.1 Orders * 4 Phylogeny
Phylogeny
* 5 References * 6 External links NAMEThe name is based upon the name " Rosidae ", which had usually been understood to be a subclass
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Taxon
In biology , a TAXON (plural TAXA; back-formation from taxonomy ) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking , especially if and when it is accepted or becomes established. It is not uncommon, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name , its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping
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Friedrich Gottlieb Bartling
FRIEDRICH GOTTLIEB BARTLING (December 9, 1798 – November 20, 1875) was a German botanist who was a native of Hanover . He studied natural sciences at the University of Göttingen , and in 1818 took a botanical journey through Hungary and Croatia . In 1822 he became a lecturer at Göttingen , where he later became a professor. In 1837 he was appointed director of its botanical garden . The plant genus Bartlingia from the family Rubiaceae is named in his honor. SELECTED PUBLICATIONS * De litoribus ac insulis maris Liburnici (1820). * Ordines naturales plantarum (1830). * Flora der österreichischen Küstenländer, (Flora of the Austrian coastal area), (1825). * Vegetabilia cellularia in Germania septentrionali praesertim in Hercynia et in agro Gottingensi lecta (1834 and 1836), with Georg Ernst Ludwig Hampe (1795–1880). The standard author abbreviation BARTL. is used to indicate this individual as the author when citing a botanical name
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Monophyly
In cladistics , a MONOPHYLETIC group is a taxon (group of organisms) which forms a clade , meaning that it consists of an ancestral species and all its descendants. Monophyletic groups are typically characterised by shared derived characteristics (synapomorphies ). The arrangement of the members of a monophyletic group is called a MONOPHYLY, synonymous with the uncommon term HOLOPHYLY. Monophyly
Monophyly
is contrasted with paraphyly and polyphyly , as shown in the second diagram. A paraphyletic group consists of all of the descendants of a common ancestor minus one or more monophyletic groups. Thus, a paraphyletic group is 'nearly' monophyletic (hence the prefix 'para', meaning 'near' or 'alongside'.) A polyphyletic group is characterized by convergent features or habits (for example, night-active primates, fruit trees, aquatic insects); the features by which the group is differentiated from others are not inherited from a common ancestor
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Cretaceous
The CRETACEOUS ( /krɪˈteɪʃəs/ , kri-TAY-shəs ) is a geologic period and system that spans 79 million years from the end of the Jurassic
Jurassic
Period 145 million years ago (mya ) to the beginning of the Paleogene Period 66 mya. It is the last period of the Mesozoic Era . The Cretaceous
Cretaceous
Period is usually abbreviated K, for its German translation Kreide (chalk). The Cretaceous
Cretaceous
was a period with a relatively warm climate , resulting in high eustatic sea levels that created numerous shallow inland seas . These oceans and seas were populated with now-extinct marine reptiles , ammonites and rudists , while dinosaurs continued to dominate on land. During this time, new groups of mammals and birds , as well as flowering plants , appeared
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Family (biology)
In biological classification , FAMILY (Latin : familia, plural familiae) is one of the eight major taxonomic ranks ; it is classified between order and genus . A family may be divided into subfamilies , which are intermediate ranks above the rank of genus . In vernacular usage , a family may be named after one of its common members; for example, walnuts and hickory trees belong to the family Juglandaceae , commonly known as the walnut family. What does or does not belong to a family—or whether a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, or any taxa. Taxonomists often take different positions about descriptions of taxa, and there may be no broad consensus across the scientific community for some time
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Clade
A CLADE (from Ancient Greek
Ancient Greek
: κλάδος, klados, "branch") is a group of organisms that consists of a common ancestor and all its lineal descendants , and represents a single "branch" on the "tree of life ". The common ancestor may be an individual, a population , a species (extinct or extant ), and so on right up to a kingdom . Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic
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Molecular Clock
The MOLECULAR CLOCK is a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged . The biomolecular data used for such calculations are usually nucleotide sequences for DNA
DNA
or amino acid sequences for proteins . The benchmarks for determining the mutation rate are often fossil or archaeological dates. The molecular clock was first tested in 1962 on the haemoglobin protein variants of various animals, and is commonly used in molecular evolution to estimate times of speciation or radiation . It is sometimes called a GENE CLOCK or an EVOLUTIONARY CLOCK
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Aptian
The APTIAN is an age in the geologic timescale or a stage in the stratigraphic column . It is a subdivision of the Early or Lower Cretaceous
Cretaceous
epoch or series and encompasses the time from 125.0 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), approximately. The Aptian succeeds the Barremian and precedes the Albian
Albian
, all part of the Lower/Early Cretaceous. The Aptian
Aptian
partly overlaps the upper part of the regionally used (in Western Europe
Western Europe
) stage Urgonian . Selli Event or OAE 1a was one of two anoxic extinction events in the Cretaceous
Cretaceous
period (in the Aptian)
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Albian
The ALBIAN is both an age of the geologic timescale and a stage in the stratigraphic column . It is the youngest or uppermost subdivision of the Early/Lower Cretaceous
Cretaceous
epoch /series . Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago). The Albian
Albian
is preceded by the Aptian and followed by the Cenomanian
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Armen Takhtajan
ARMEN LEONOVICH TAKHTAJAN or TAKHTAJIAN (Armenian : Արմեն Լևոնի Թախտաջյան; Russian : Армен Леонович Тахтаджян; surname also transliterated Takhtadjan, Takhtadzhi︠a︡n or Takhtadzhian, pronounced TAHK-tuh-jahn) (June 10, 1910 – November 13, 2009), was a Soviet -Armenian botanist , one of the most important figures in 20th century plant evolution and systematics and biogeography . His other interests included morphology of flowering plants , paleobotany , and the flora of the Caucasus
Caucasus
. He was born in Shusha . He was one of the most influential taxonomists of the latter twentieth century
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ICBN
The INTERNATIONAL CODE OF NOMENCLATURE FOR ALGAE, FUNGI, AND PLANTS (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all those "traditionally treated as algae, fungi, or plants". :Preamble, para. 8 It was formerly called the INTERNATIONAL CODE OF BOTANICAL NOMENCLATURE (ICBN); the name was changed at the International Botanical Congress in Melbourne in July 2011 as part of the Melbourne Code which replaces the Vienna Code of 2005. As with previous codes, it took effect as soon as it was ratified by the congress (on Saturday 23 July 2011), but the documentation of the code in its final form was not finished until some time after the congressional meeting. Preliminary wording of some of the articles with the most significant changes has been published in September 2011. The name of the Code is partly capitalized and partly not
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Taxonomic Rank
In biological classification , TAXONOMIC RANK is the relative level of a group of organisms (a taxon ) in a taxonomic hierarchy . Examples of taxonomic ranks are species , genus , family , order , class , phylum , kingdom , domain , etc. A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of traits or features from common ancestors. The rank of any species and the description of its genus is basic; which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two. Consider a particular species, the red fox , Vulpes
Vulpes
vulpes: its next rank, the genus Vulpes
Vulpes
, comprises all the 'true foxes'
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Monophyletic
In cladistics , a MONOPHYLETIC group is a taxon (group of organisms) which forms a clade , meaning that it consists of an ancestral species and all its descendants. Monophyletic groups are typically characterised by shared derived characteristics (synapomorphies ). The arrangement of the members of a monophyletic group is called a MONOPHYLY, synonymous with the uncommon term HOLOPHYLY. Monophyly is contrasted with paraphyly and polyphyly , as shown in the second diagram. A paraphyletic group consists of all of the descendants of a common ancestor minus one or more monophyletic groups. Thus, a paraphyletic group is 'nearly' monophyletic (hence the prefix 'para', meaning 'near' or 'alongside'.) A polyphyletic group is characterized by convergent features or habits (for example, night-active primates, fruit trees, aquatic insects); the features by which the group is differentiated from others are not inherited from a common ancestor
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Circumscription (taxonomy)
In biological taxonomy , CIRCUMSCRIPTION is the definition of a taxon , that is, a group of organisms . One goal of biological taxonomy is to achieve a stable circumscription for every taxon. Achieving stability is not yet a certainty in most taxa, and many that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics . In essence, new discoveries may invalidate the application of irrelevant attributes used in established or obsolete circumscriptions, or present new attributes useful in cladistic taxonomy . An example of a taxonomic group with unstable circumscription is Anacardiaceae , a family of flowering plants . Some experts favor a circumscription in which this family includes the Blepharocaryaceae , Julianaceae , and Podoaceae , which are sometimes considered to be separate families
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Biological Classification
TAXONOMY (from Ancient Greek τάξις (taxis ), meaning 'arrangement', and -νομία (-nomia), meaning 'method ') is the science of defining and naming groups of biological organisms on the basis of shared characteristics. Organisms are grouped together into taxa (singular: taxon) and these groups are given a taxonomic rank ; groups of a given rank can be aggregated to form a super group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are kingdom, phylum, class, order, family, genus and species. The Swedish botanist Carl Linnaeus
Carl Linnaeus
is regarded as the father of taxonomy, as he developed a system known as Linnaean taxonomy for categorization of organisms and binomial nomenclature for naming organisms
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