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Nymphaeales
The Nymphaeales are an order of flowering plants, consisting of three families of aquatic plants, the Hydatellaceae, the Cabombaceae, and the Nymphaeaceae (water lilies). It is one of the three orders of basal angiosperms, an early-diverging grade of flowering plants. At least 10 morphological characters unite the Nymphaeales.Peter F. Stevens. 2001 onwards. Angiosperm Phylogeny Website At: Missouri Botanical Garden Website. (see ''External links'' below). One of the traits is the absence of a vascular cambium, which is required to produce both xylem (wood) and phloem, which therefore are missing. Molecular synapomorphies are also known. The Plant List, created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden recognizes about 70 species in 11 genera within the order,The Plant List (website). 2010. (See ''External links'' below.) but a phylogenetic study of the genus ''Nymphaea'' implies that the number of species could be more than 90. The difference in species nu ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in ...
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Basal Angiosperms
The basal angiosperms are the flowering plants which diverged from the lineage leading to most flowering plants. In particular, the most basal angiosperms were called the ANITA grade, which is made up of ''Amborella'' (a single species of shrub from New Caledonia), Nymphaeales (water lilies, together with some other aquatic plants) and Austrobaileyales (woody aromatic plants including star anise). ''ANITA'' stands for ''Amborella'', ''N''ymphaeales, ''I'' lliciales, ''T''rimeniaceae, and '' Austrobaileya''. Some authors have shortened this to ''ANA''-grade for the three orders, ''A''mborellales, ''N''ymphaeales, and ''A''ustrobaileyales, since the order Iliciales was reduced to the family Illiciaceae and placed, along with the family Trimeniaceae, within the Austrobaileyales. The basal angiosperms are only a few hundred species, compared with hundreds of thousands of species of eudicots, monocots, and magnoliids. They diverged from the ancestral angiosperm lineage befo ...
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Nymphaeaceae
Nymphaeaceae () is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate and tropical climates around the world. The family contains nine genera with about 70 known species. Water lilies are rooted in soil in bodies of water, with leaves and flowers floating on or emergent from the surface. Leaves are round, with a radial notch in '' Nymphaea'' and '' Nuphar'', but fully circular in '' Victoria'' and '' Euryale''. Water lilies are a well-studied clade of plants because their large flowers with multiple unspecialized parts were initially considered to represent the floral pattern of the earliest flowering plants, and later genetic studies confirmed their evolutionary position as basal angiosperms. Analyses of floral morphology and molecular characteristics and comparisons with a sister taxon, the family Cabombaceae, indicate, however, that the flowers of extant water lilies with the most floral parts are more d ...
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Cabombaceae
The Cabombaceae are a family of aquatic, herbaceous flowering plants. A common name for its species is water shield. The family is recognised as distinct in the Angiosperm Phylogeny Group IV system (2016). The family consists of two genera of aquatic plants, '' Brasenia'' and ''Cabomba'', totalling six species. The Cabombaceae are all aquatic, living in still or slow-moving waters of temperate and tropical North and South America, Europe, Asia, Africa, and Australia. Although found on all continents but Antarctica, the plants tend to grow in relatively restricted ranges. The family has an extensive fossil record from the Cretaceous with plants that exhibit affinities to either the Cabombaceae or Nymphaeaceae occurring in the Early Cretaceous. One such likely Cretaceous member is the genus '' Pluricarpellatia'', found in rocks 115 million years old in what is now Brazil. The APG system of 1998 included this family in the water lily family Nymphaeaceae, as did the APG II syste ...
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Hydatellaceae
Hydatellaceae are a family of small, aquatic flowering plants. The family consists of tiny, relatively simple plants occurring in Australasia and India. It was formerly considered to be related to the grasses and sedges (order Poales), but has been reassigned to the order Nymphaeales as a result of DNA and morphological analyses showing that it represents one of the earliest groups to split off in flowering-plant phylogeny, rather than having a close relationship to monocots, which it bears a superficial resemblance to due to convergent evolution. The family includes only the genus '' Trithuria'', which has at least 13 species, although species diversity in the family has probably been substantially underestimated. Description Plants are submerged and emergent aquatic plants, rooted in the substrate below the water. They are tiny plants, just a few cm tall. Most species are ephemeral aquatics that flower in vernal pools when the water draws down, but several species are submer ...
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Nymphaea
''Nymphaea'' () is a genus of hardy and tender aquatic plants in the family Nymphaeaceae. The genus has a cosmopolitan distribution. Many species are cultivated as ornamental plants, and many cultivars have been bred. Some taxa occur as introduced species where they are not native,''Nymphaea''.
Flora of North America.
and some are weeds.''Nymphaea''.
The Jepson eFlora 2013.
Plants of the genus are known commonly as water lilies,
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Vascular Cambium
The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark. In herbaceous plants, it occurs in the vascular bundles which are often arranged like beads on a necklace forming an interrupted ring inside the stem. In woody plants, it forms a cylinder of unspecialized meristem cells, as a continuous ring from which the new tissues are grown. Unlike the xylem and phloem, it does not transport water, minerals or food through the plant. Other names for the vascular cambium are the main cambium, wood cambium, or bifacial cambium. Occurrence Vascular cambia are found in all seed plants except for five angiosperm lineages which have independently lost it; Nymphaeales, Ceratophyllum, Nelumbo, Podostemaceae, and monoco ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and diversification. However, such a correlation does not make a given case predicable, so ancestral characters should not be imputed to ...
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Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Megaannum#SI prefix multipliers, Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining Anoxic event#Cretaceous, early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java Plateau, Ontong Java-Manihiki Plateau, Manihiki-Hikurangi Plateau, Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. T ...
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The Plant List
The Plant List was a list of botanical names of species of plants created by the Royal Botanic Gardens, Kew and the Missouri Botanical Garden and launched in 2010. It was intended to be a comprehensive record of all known names of plant species over time, and was produced in response to Target 1 of the 2002-2010 Global Strategy for Plant Conservation (GSP C), to produce "An online flora of all known plants.” It has not been updated since 2013, and has been superseded by World Flora Online. World Flora Online In October 2012, the follow-up project World Flora Online was launched with the aim to publish an online flora of all known plants by 2020. This is a project of the United Nations Convention on Biological Diversity, with the aim of halting the loss of plant species worldwide by 2020. It is developed by a collaborative group of institutions around the world response to the 2011-2020 GSPC's updated Target 1. This aims to achieve an online Flora of all known plants by 202 ...
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Royal Botanic Gardens, Kew
Royal Botanic Gardens, Kew is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. An internationally important botanical research and education institution, it employs 1,100 staff. Its board of trustees is chaired by Dame Amelia Fawcett. The organisation manages botanic gardens at Kew in Richmond upon Thames in south-west London, and at Wakehurst, a National Trust property in Sussex which is home to the internationally important Millennium Seed Bank, whose scientists work with partner organisations in more than 95 countries. Kew, jointly with the Forestry Commission, founded Bedgebury National Pinetum in Kent in 1923, specialising in growing conifers. In 1994, the Castle Howard Arboretum Trust, which runs the Yorkshire Arboretum, was formed as a partnership between Kew and the Castle Howard Estate. In 2019, the organisation had 2,316,699 public visitors at Kew, and 312,813 at Wakehurst. Its site at ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and do ...
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