Hydatellaceae
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Hydatellaceae
Hydatellaceae are a family of small, aquatic flowering plants. The family consists of tiny, relatively simple plants occurring in Australasia and India. It was formerly considered to be related to the grasses and sedges (order Poales), but has been reassigned to the order Nymphaeales as a result of DNA and morphological analyses showing that it represents one of the earliest groups to split off in flowering-plant phylogeny, rather than having a close relationship to monocots, which it bears a superficial resemblance to due to convergent evolution. The family includes only the genus '' Trithuria'', which has at least 13 species, although species diversity in the family has probably been substantially underestimated. Description Plants are submerged and emergent aquatic plants, rooted in the substrate below the water. They are tiny plants, just a few cm tall. Most species are ephemeral aquatics that flower in vernal pools when the water draws down, but several species are submer ...
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Nymphaeales
The Nymphaeales are an order of flowering plants, consisting of three families of aquatic plants, the Hydatellaceae, the Cabombaceae, and the Nymphaeaceae (water lilies). It is one of the three orders of basal angiosperms, an early-diverging grade of flowering plants. At least 10 morphological characters unite the Nymphaeales.Peter F. Stevens. 2001 onwards. Angiosperm Phylogeny Website At: Missouri Botanical Garden Website. (see ''External links'' below). One of the traits is the absence of a vascular cambium, which is required to produce both xylem (wood) and phloem, which therefore are missing. Molecular synapomorphies are also known. The Plant List, created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden recognizes about 70 species in 11 genera within the order,The Plant List (website). 2010. (See ''External links'' below.) but a phylogenetic study of the genus '' Nymphaea'' implies that the number of species could be more than 90. The difference in spec ...
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Trithuria Submersa In Fruit
''Trithuria'' is a genus of small aquatic herb, which represent the only members of the family Hydatellaceae found in India, Australia, and New Zealand. Most of the 12 formally characterised species of ''Trithuria'' are found in Australia, with the exception of ''T. inconspicua'' and ''T. konkanensis'', which are found in New Zealand and India, respectively.Dmitry D. Sokoloff, Margarita V. Remizowa, Terry D. Macfarlane, and Paula J. Rudall. 2008. "Classification of the early-divergent angiosperm family Hydatellaceae: one genus instead of two, four new species and sexual dimorphism in dioecious taxa". ''Taxon'' 57(1):179-200.Yadav SR, Janarthanam MK. 1995 Trithuria konkanensis (Hydatellaceae), eine neue Art aus Indien. ''Aqua Planta'' 20. (3): 91-97 (1995). Until genetic testing proved otherwise, these plants were believed to be Monocots related to the grasses (Poaceae). They are unique in being the only plants besides Lacandonia schizmatica and L. braziliana in which the stamen ...
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Trithuria
''Trithuria'' is a genus of small aquatic herb, which represent the only members of the family Hydatellaceae found in India, Australia, and New Zealand. Most of the 12 formally characterised species of ''Trithuria'' are found in Australia, with the exception of ''T. inconspicua'' and ''T. konkanensis'', which are found in New Zealand and India, respectively.Dmitry D. Sokoloff, Margarita V. Remizowa, Terry D. Macfarlane, and Paula J. Rudall. 2008. "Classification of the early-divergent angiosperm family Hydatellaceae: one genus instead of two, four new species and sexual dimorphism in dioecious taxa". ''Taxon'' 57(1):179-200.Yadav SR, Janarthanam MK. 1995 Trithuria konkanensis (Hydatellaceae), eine neue Art aus Indien. ''Aqua Planta'' 20. (3): 91-97 (1995). Until genetic testing proved otherwise, these plants were believed to be Monocots related to the grasses (Poaceae). They are unique in being the only plants besides Lacandonia schizmatica and L. braziliana in which the stamens ...
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Cabombaceae
The Cabombaceae are a family of aquatic, herbaceous flowering plants. A common name for its species is water shield. The family is recognised as distinct in the Angiosperm Phylogeny Group IV system (2016). The family consists of two genera of aquatic plants, '' Brasenia'' and ''Cabomba'', totalling six species. The Cabombaceae are all aquatic, living in still or slow-moving waters of temperate and tropical North and South America, Europe, Asia, Africa, and Australia. Although found on all continents but Antarctica, the plants tend to grow in relatively restricted ranges. The family has an extensive fossil record from the Cretaceous with plants that exhibit affinities to either the Cabombaceae or Nymphaeaceae occurring in the Early Cretaceous. One such likely Cretaceous member is the genus '' Pluricarpellatia'', found in rocks 115 million years old in what is now Brazil. The APG system of 1998 included this family in the water lily family Nymphaeaceae, as did the APG II syste ...
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Pseudanthium
A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, composite flowers, or capitula, which are special types of inflorescences in which anything from a small cluster to hundreds or sometimes thousands of flowers are grouped together to form a single flower-like structure. Pseudanthia take various forms. The real flowers (the florets) are generally small and often greatly reduced, but the pseudanthium itself can sometimes be quite large (as in the heads of some varieties of sunflower). Pseudanthia are characteristic of the daisy and sunflower family (Asteraceae), whose flowers are differentiated into ray flowers and disk flowers, unique to this family. The disk flowers in the center of the pseudanthium are actinomorphic and the corolla is fused into a tube. Flowers on the periphery are zygomor ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils ar ...
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Poales
The Poales are a large order of flowering plants in the monocotyledons, and includes families of plants such as the grasses, bromeliads, and sedges. Sixteen plant families are currently recognized by botanists to be part of Poales. Description The flowers are typically small, enclosed by bracts, and arranged in inflorescences (except in three species of the genus '' Mayaca'', which possess very reduced, one-flowered inflorescences). The flowers of many species are wind pollinated; the seeds usually contain starch. Taxonomy The APG III system (2009) accepts the order within a monocot clade called commelinids, and accepts the following 16 families: The earlier APG system (1998) adopted the same placement of the order, although it used the spelling "commelinoids". It did not include the Bromeliaceae and Mayaceae, but had the additional families Prioniaceae (now included in Thurniaceae), Sparganiaceae (now in Typhaceae), and Hydatellaceae (now transferred out of the monocots; ...
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APG IV System
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one. Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that ar ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families follo ...
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Botanical Journal Of The Linnean Society
The ''Botanical Journal of the Linnean Society'' is a scientific journal publishing original papers relating to the taxonomy of all plant groups and fungi, including anatomy, biosystematics, cytology, ecology, ethnobotany, electron microscopy, morphogenesis, palaeobotany, palynology and phytochemistry.Botanical Journal of the Linnean Society
The journal is published by the and is available in both print and searchable online formats. Like the ''

Derived Trait
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis ...
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Nymphaeaceae
Nymphaeaceae () is a family of flowering plants, commonly called water lilies. They live as rhizomatous aquatic herbs in temperate and tropical climates around the world. The family contains nine genera with about 70 known species. Water lilies are rooted in soil in bodies of water, with leaves and flowers floating on or emergent from the surface. Leaves are round, with a radial notch in '' Nymphaea'' and '' Nuphar'', but fully circular in '' Victoria'' and '' Euryale''. Water lilies are a well-studied clade of plants because their large flowers with multiple unspecialized parts were initially considered to represent the floral pattern of the earliest flowering plants, and later genetic studies confirmed their evolutionary position as basal angiosperms. Analyses of floral morphology and molecular characteristics and comparisons with a sister taxon, the family Cabombaceae, indicate, however, that the flowers of extant water lilies with the most floral parts are more d ...
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