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Clitellata
The Clitellata are a class of annelid worms, characterized by having a clitellum - the 'collar' that forms a reproductive cocoon during part of their life cycles. The clitellates comprise around 8,000 species. Unlike the class of Polychaeta, they do not have parapodia and their heads are less developed. Characteristics Clitellate annelids are segmented worms characterised by the clitellum or girdle which is located near the head end of mature individuals. The mouth is on the ventral surface and is overhung by the prostomium (proboscis). The brain is not located in the head but in one of the body segments. The clitellum is formed by a modification of several segments, and either includes the female gonopores or is located just behind them. During copulation, this glandular tissue secretes mucus that keeps the paired individuals together while they exchange sperm. Afterwards it secretes material that forms a cocoon that encircles the animal's body and encloses the eggs and sp ...
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Annelid
The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The Annelids are bilaterally symmetrical, triploblastic, coelomate, invertebrate organisms. They also have parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-groups of polychae ...
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Hirudinea
Leeches are segmented parasitic or predatory worms that comprise the subclass Hirudinea within the phylum Annelida. They are closely related to the oligochaetes, which include the earthworm, and like them have soft, muscular segmented bodies that can lengthen and contract. Both groups are hermaphrodites and have a clitellum, but leeches typically differ from the oligochaetes in having suckers at both ends and in having ring markings that do not correspond with their internal segmentation. The body is muscular and relatively solid, and the coelom, the spacious body cavity found in other annelids, is reduced to small channels. The majority of leeches live in freshwater habitats, while some species can be found in terrestrial or marine environments. The best-known species, such as the medicinal leech, ''Hirudo medicinalis'', are hematophagous, attaching themselves to a host with a sucker and feeding on blood, having first secreted the peptide hirudin to prevent the blood from ...
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Leech
Leeches are segmented parasitism, parasitic or Predation, predatory worms that comprise the Class (biology), subclass Hirudinea within the phylum Annelida. They are closely related to the Oligochaeta, oligochaetes, which include the earthworm, and like them have soft, muscular segmented bodies that can lengthen and contract. Both groups are hermaphrodites and have a clitellum, but leeches typically differ from the oligochaetes in having suckers at both ends and in having ring markings that do not correspond with their internal segmentation. The body is muscular and relatively solid, and the coelom, the spacious body cavity found in other annelids, is reduced to small channels. The majority of leeches live in freshwater habitats, while some species can be found in terrestrial or marine environments. The best-known species, such as the medicinal leech, ''Hirudo medicinalis'', are hematophagous, attaching themselves to a host with a sucker and feeding on blood, having first secr ...
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Branchiobdellida
Branchiobdellida is an order of freshwater leech-like clitellates that are obligate Ectosymbiosis, ectosymbionts or ectoparasites, mostly of astacoidean crayfish. They are found in the Northern Hemisphere and have a Holarctic realm, holarctic distribution in East Asia, the Euro-Mediterranean region and North and Central America, with the greatest species diversity being in North and Central America. Taxonomy The exact relationship of this group of clitellates with other groups has been the subject of debate for a century or more. The American zoologist Perry C. Holt wrote his doctoral thesis on branchiobdellids and devoted his 45 years of research to the taxon. With his colleague, the American zoologist Richard L. Hoffman, he described 8 new Genus, genera and 75 new species, and re-described various other species. In 1965, he came to the conclusion that the group should be raised from a Family (biology), family to an Order (biology), order, a sister group to the Oligochaeta, oligo ...
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Branchiobdellae
Branchiobdellida is an order of freshwater leech-like clitellates that are obligate ectosymbionts or ectoparasites, mostly of astacoidean crayfish. They are found in the Northern Hemisphere and have a holarctic distribution in East Asia, the Euro-Mediterranean region and North and Central America, with the greatest species diversity being in North and Central America. Taxonomy The exact relationship of this group of clitellates with other groups has been the subject of debate for a century or more. The American zoologist Perry C. Holt wrote his doctoral thesis on branchiobdellids and devoted his 45 years of research to the taxon. With his colleague, the American zoologist Richard L. Hoffman, he described 8 new genera and 75 new species, and re-described various other species. In 1965, he came to the conclusion that the group should be raised from a family to an order, a sister group to the oligochaetes and leeches. Forty years later, molecular data from rDNA and mitochon ...
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Oligochaeta
Oligochaeta () is a subclass of animals in the phylum Annelida, which is made up of many types of aquatic and terrestrial worms, including all of the various earthworms. Specifically, oligochaetes comprise the terrestrial megadrile earthworms (some of which are semiaquatic or fully aquatic), and freshwater or semiterrestrial microdrile forms, including the tubificids, pot worms and ice worms ( Enchytraeidae), blackworms ( Lumbriculidae) and several interstitial marine worms. With around 10,000 known species, the Oligochaeta make up about half of the phylum Annelida. These worms usually have few setae (chaetae) or "bristles" on their outer body surfaces, and lack parapodia, unlike polychaeta. Diversity Oligochaetes are well-segmented worms and most have a spacious body cavity (coelom) used as a hydroskeleton. They range in length from less than up to in the 'giant' species such as the giant Gippsland earthworm (''Megascolides australis'') and the Mekong worm (''Amynth ...
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Clitellum
The clitellum is a thickened glandular and non-segmented section of the body wall near the head in earthworms and leeches, that secretes a viscid sac in which eggs are stored. It is located near the anterior end of the body, between the fourteenth and seventeenth segments. The number of the segments to where the clitellum begins and the number of segments that make up the clitellum are important for identifying earthworms. In microdrile earthworms, the clitellum has only one layer, resulting in a smaller quantity of eggs than that of the megadrile earthworms, which have larger multi-layered clitellum that have special cells that secrete albumin into the worms' egg sac. The clitellum is part of the reproductive system of clitellates, a subgroup of annelids which contains oligochaetes (earthworms) and hirudineans (leeches). The clitellum is a thick, saddle-like ring found in the epidermis (skin) of the worm, usually with a light-colored pigment. To form a cocoon for its eggs, t ...
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Hermaphrodite
In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have separate sexes. In these groups, hermaphroditism is a normal condition, enabling a form of sexual reproduction in which either partner can act as the female or male. For example, the great majority of tunicates, pulmonate molluscs, opisthobranch, earthworms, and slugs are hermaphrodites. Hermaphroditism is also found in some fish species and to a lesser degree in other vertebrates. Most plants are also hermaphrodites. Animal species having different sexes, male and female, are called gonochoric, which is the opposite of hermaphrodite. There are also species where hermaphrodites exist alongside males (called androdioecy) or alongside females (called gynodioecy), or all three exist in the same species (called trioecy); these three syst ...
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Zygote
A zygote (, ) is a eukaryotic cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individual organism. In multicellular organisms, the zygote is the earliest developmental stage. In humans and most other anisogamous organisms, a zygote is formed when an egg cell and sperm cell come together to create a new unique organism. In single-celled organisms, the zygote can divide asexually by mitosis to produce identical offspring. German zoologists Oscar and Richard Hertwig made some of the first discoveries on animal zygote formation in the late 19th century. Humans In human fertilization, a released ovum (a haploid secondary oocyte with replicate chromosome copies) and a haploid sperm cell ( male gamete) combine to form a single diploid cell called the zygote. Once the single sperm fuses with the oocyte, the latter completes the division of the se ...
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Ovum
The egg cell, or ovum (plural ova), is the female reproductive cell, or gamete, in most anisogamous organisms (organisms that reproduce sexually with a larger, female gamete and a smaller, male one). The term is used when the female gamete is not capable of movement (non- motile). If the male gamete ( sperm) is capable of movement, the type of sexual reproduction is also classified as oogamous. A nonmotile female gamete formed in the oogonium of some algae, fungi, oomycetes, or bryophytes is an oosphere. When fertilized the oosphere becomes the oospore. When egg and sperm fuse during fertilisation, a diploid cell (the zygote) is formed, which rapidly grows into a new organism. History While the non-mammalian animal egg was obvious, the doctrine ''ex ovo omne vivum'' ("every living nimal comes froman egg"), associated with William Harvey (1578–1657), was a rejection of spontaneous generation and preformationism as well as a bold assumption that mammals also reprodu ...
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Ovary
The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. The ovaries also secrete hormones that play a role in the menstrual cycle and fertility. The ovary progresses through many stages beginning in the prenatal period through menopause. It is also an endocrine gland because of the various hormones that it secretes. Structure The ovaries are considered the female gonads. Each ovary is whitish in color and located alongside the lateral wall of the uterus in a region called the ovarian fossa. The ovarian fossa is the region that is bounded by the external iliac artery and in front of the ureter and the internal iliac artery. This area is about 4 cm x 3 cm x 2 cm in size.Daftary, Shirish; Chakravarti, Sudip (2011). Manual of Obstetrics, 3rd Edition. Elsevier. pp. 1-16. . The ...
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Thymallus
''Thymallus'' is a genus of freshwater fish in the salmon family Salmonidae; it is the only genus of subfamily Thymallinae. The type species is '' Thymallus thymallus,'' the grayling. The species in the genus are generically called graylings, but without qualification this also refers specifically to ''T. thymallus.'' Distribution The fishes of this genus are native to the northern parts of the Palearctic and Nearctic realms, ranging from the United Kingdom and northern Europe across Eurasia to Siberia, as well as northern North America. ''T. thymallus'', the grayling, is widespread in Europe, and ''T. arcticus,'' the Arctic grayling, is widespread throughout Eurasia east of the Ural Mountains and in the Nearctic. The other species have more localized ranges in northern Asia. Appearance ''Thymallus'' species are distinguished from other members of the salmon family by their larger scales, their small mouths with teeth on the maxillary bone, and most striking of all, their show ...
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