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The yellow-headed caracara (''Daptrius chimachima'') is a
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
. It is found in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north and ...
South America and the southern portion of
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. C ...
. Unlike the
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
s in the same family, the caracara is not a fast-flying aerial hunter, but is rather sluggish and often obtains food by
scavenging Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding be ...
.


Taxonomy

When
Louis Jean Pierre Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collecte ...
first described the yellow-headed caracara in 1816, he gave it the scientific name ''Polyborus chimachima'', putting it in the same genus as the crested caracaras. In 1824, German naturalist
Johann Baptist von Spix Johann Baptist Ritter von Spix (9 February 1781 – 13 March 1826) was a German biologist. From his expedition to Brazil, he brought to Germany a large variety of specimens of plants, insects, mammals, birds, amphibians and fish. They constitute ...
created the genus ''Milvago'' for this species and the closely related
chimango caracara The chimango caracara (''Milvago chimango'') is a species of bird of prey in the family Falconidae. It is found in Argentina, Uruguay, Chile, Paraguay and south of Brazil. The chimango is found as far south as Tierra del Fuego and is a vagrant to ...
. Formerly, the yellow-headed caracara was classified in the genus ''Milvago'' together with the
Chimango caracara The chimango caracara (''Milvago chimango'') is a species of bird of prey in the family Falconidae. It is found in Argentina, Uruguay, Chile, Paraguay and south of Brazil. The chimango is found as far south as Tierra del Fuego and is a vagrant to ...
(then ''Milvago chimango''). However, molecular studies from 2012 showed that the genus ''Milvago'' is not
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
. These two species were therefore reclassified together with various other species in the genus ''Phalcoboenus'' with the species' closest relative (the
Black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guiana lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republic of Suriname ...
, ''Daptrius ater''). These species are now all part of the genus ''Daptrius''. These changes have been adopted in the 2022 edition of
The Clements Checklist of Birds of the World ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
.


Description

The yellow-headed caracara is in length and weighs on average. Like many other birds of prey, the female is larger than the male, weighing against the male's . Apart from the difference in size, there is no significant
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most anim ...
in this species. It is broad-winged and long-tailed, somewhat resembling a small ''
Buteo ''Buteo'' is a genus of medium to fairly large, wide-ranging raptors with a robust body and broad wings. In the Old World, members of this genus are called "buzzards", but "hawk" is used in the New World (Etymology: ''Buteo'' is the Latin name o ...
''. The adult has a buff head, with a black streak behind the eye, and buff underparts. The upper
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
is brown with distinctive pale patches on the
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s of the wings, and the tail is barred cream and brown. The head and underparts of immature birds have dense brown mottling. The voice is a characteristic screamed '.


Distribution and habitat

This is a bird of
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s and
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
edges. Resident from Costa Rica south through
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of Gr ...
to northern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
(the provinces of
Misiones Misiones (, ''Missions'') is one of the 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamia region. It is surrounded by Paraguay to the northwest, Brazil to the north, east and south, and Corrientes ...
, Chaco,
Formosa Taiwan, officially the Republic of China (ROC), is an island country located in East Asia. The main island of Taiwan, formerly known in the Western political circles, press and literature as Formosa, makes up 99% of the land area of the territori ...
,
Corrientes Corrientes (; Guaraní: Taragüí, literally: "Currents") is the capital city of the province of Corrientes, Argentina, located on the eastern shore of the Paraná River, about from Buenos Aires and from Posadas, on National Route 12. It ha ...
and Santa Fe), it is typically found from sea level to , and occasionally to
above mean sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...
. In southern South America, it is replaced by a close relative, the
chimango caracara The chimango caracara (''Milvago chimango'') is a species of bird of prey in the family Falconidae. It is found in Argentina, Uruguay, Chile, Paraguay and south of Brazil. The chimango is found as far south as Tierra del Fuego and is a vagrant to ...
, whose range overlaps with that of the yellow-headed caracara in southern Brazil, northern Argentina,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to ...
and
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
. A larger and stouter
paleosubspecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
, ''Milvago chimachima readei'', occurred in
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to th ...
and possibly elsewhere some tens of thousand years ago, during the
Late Pleistocene The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within ...
. According to the Peregrine Fund database, the yellow-headed caracara is expanding its range into
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
.


Behavior and ecology

The yellow-headed caracara is
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
, and will eat
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
s,
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
,
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates ( lizards and snakes) and rhynchoce ...
s,
amphibian Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arborea ...
s,
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s and other small animals, as well as
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
, dung and
garbage Garbage, trash, rubbish, or refuse is waste material that is discarded by humans, usually due to a perceived lack of utility. The term generally does not encompass bodily waste products, purely liquid or gaseous wastes, or toxic waste produc ...
. Birds (such as nestlings and young chickens) are rarely if ever taken, and this species will not elicit warning calls from
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These are ...
s that cross its path even in open
cerrado The ''Cerrado'' (, ) is a vast ecoregion of tropical savanna in eastern Brazil, particularly in the states of Goiás, Mato Grosso do Sul, Mato Grosso, Tocantins, Minas Gerais, and the Federal District. The core areas of the Cerrado biome are th ...
habitat. It will also remove and eat
tick Ticks (order Ixodida) are parasitic arachnids that are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, species, and "fullness". Ticks are external parasites, living by ...
s from
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
and Baird's tapir (''Tapirus bairdii''), and is sometimes called the ''tickbird''. It has been observed also to forage for small invertebrates in the fur of
brown-throated three-toed sloth The brown-throated sloth (''Bradypus variegatus'') is a species of three-toed sloth found in the Neotropical realm of Central and South America. It is the most common of the four species of three-toed sloth, and is found in the forests of Sout ...
s and
capybara The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydro ...
s. They feed not only on adult
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of ...
s but also on pupae and larvae of
Lepidoptera Lepidoptera ( ) is an order of insects that includes butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 percent of the total described specie ...
,
Diptera Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced ...
and
Orthoptera Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshop ...
. In addition, at least younger birds are fond of certain
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particul ...
s, such as those of the
coconut palm The coconut tree (''Cocos nucifera'') is a member of the palm tree family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the f ...
(''Cocos nucifera''),
oil palm ''Elaeis'' () is a genus of palms containing two species, called oil palms. They are used in commercial agriculture in the production of palm oil. The African oil palm ''Elaeis guineensis'' (the species name ''guineensis'' referring to its cou ...
(''Elaeis guineensis'') and pequi (''
Caryocar brasiliense ''Caryocar brasiliense'', known as pequi (, ) or "souari nut", like its congeners, is an edible fruit popular in some areas of Brazil, especially in Centerwestern Brazil. Taxonomy The pequi tree grows up to 10 m (30 ft) tall. It i ...
''). It lays from five to seven brown-marked buff
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
in a stick
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
in a tree. The yellow-headed caracara has benefited from forest clearing for cattle ranching. Its status in
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmo ...
has changed from rare to fairly common, and it was first seen on
Tobago Tobago () is an island and ward within the Republic of Trinidad and Tobago. It is located northeast of the larger island of Trinidad and about off the northeastern coast of Venezuela. It also lies to the southeast of Grenada. The official b ...
in 1987. It adapts readily to
urban areas An urban area, built-up area or urban agglomeration is a human settlement with a high population density and infrastructure of built environment. Urban areas are created through urbanization and are categorized by urban morphology as cities, ...
and, together with species such as the American black vulture (''Coragyps atratus''), it is among the most commonly seen bird of prey in
Latin America Latin America or * french: Amérique Latine, link=no * ht, Amerik Latin, link=no * pt, América Latina, link=no, name=a, sometimes referred to as LatAm is a large cultural region in the Americas where Romance languages — languages derived f ...
n cities. Consequently, this wide-ranging species has been assigned a risk level of
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
category on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
. In Panama City for example, as a result of the increased urban sprawl, yellow-headed caracara pairs are frequently seen along the rooftops in suburban neighborhoods.


Gallery

Image:Yellow-headed Caracara RWD2.jpg, Adult in
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cost ...
Image:Yellow-headed Caracara RWD3.jpg, Juvenile in
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cost ...
Image: Yellow-headed caracara (Milvago chimachima) on capybara (Hydrochoeris hydrochaeris).JPG, Adult on capybara (''Hydrochoeris hydrochaeris''), the
Pantanal The Pantanal () is a natural region encompassing the world's largest tropical wetland area, and the world's largest flooded grasslands. It is located mostly within the Brazilian state of Mato Grosso do Sul, but it extends into Mato Grosso and p ...
, Brazil File:Milvago chimachima flying.jpg, Among the most commonly seen birds of prey in Latin American cities File:Flickr - Dario Sanches - CARRAPATEIRO ( Milvago chimachima ).jpg


References


Further reading

*


External links

* *
Yellow-headed caracara Photo
a
panama-guide.com
* * {{Taxonbar, from=Q28484
yellow-headed caracara The yellow-headed caracara (''Daptrius chimachima'') is a bird of prey in the family Falconidae. It is found in tropical and subtropical South America and the southern portion of Central America. Unlike the falcons in the same family, the carac ...
yellow-headed caracara The yellow-headed caracara (''Daptrius chimachima'') is a bird of prey in the family Falconidae. It is found in tropical and subtropical South America and the southern portion of Central America. Unlike the falcons in the same family, the carac ...
Birds of South America Birds of Costa Rica Birds of Panama Birds of Trinidad and Tobago Birds of the Caribbean
yellow-headed caracara The yellow-headed caracara (''Daptrius chimachima'') is a bird of prey in the family Falconidae. It is found in tropical and subtropical South America and the southern portion of Central America. Unlike the falcons in the same family, the carac ...
yellow-headed caracara The yellow-headed caracara (''Daptrius chimachima'') is a bird of prey in the family Falconidae. It is found in tropical and subtropical South America and the southern portion of Central America. Unlike the falcons in the same family, the carac ...