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''Quercus lobata'', commonly called the valley oak or roble, grows into the largest of California
oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ...
s. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to California, growing in interior valleys and foothills from
Siskiyou County Siskiyou County (, ) is a county in the northernmost part of the U.S. state of California. As of the 2020 census, the population was 44,076. Its county seat is Yreka and its highest point is Mount Shasta. It falls within the Cascadia bioregio ...
to
San Diego County San Diego County (), officially the County of San Diego, is a county in the southwestern corner of the U.S. state of California. As of the 2020 census, the population was 3,298,634, making it California's second-most populous county and the f ...
. Mature specimens may attain an age of up to 600 years. This
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
oak requires year-round access to groundwater. Its thick, ridged bark is characteristic and resembles
alligator An alligator is a large reptile in the Crocodilia order in the genus ''Alligator'' of the family Alligatoridae. The two extant species are the American alligator (''A. mississippiensis'') and the Chinese alligator (''A. sinensis''). Additional ...
hide. The valley oak's deeply lobed leaves assist in identification.


Description

The valley oak may surpass in height, with a sturdy trunk possibly exceeding in diameter. The "Henley Oak", in
Covelo, California Covelo ( ) is a census-designated place (CDP) in Mendocino County, California, United States. Covelo is located as the crow flies east-northeast of Laytonville, at an elevation of . The population was 1,394 at the 2020 census, up from 1,255 at ...
, is the tallest known valley oak, at . The branches have an irregular, spreading and arching appearance that produce a profound leafless silhouette in the clear winter sky. During autumn, the
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
turn a yellow to light orange color but become brown later in the season. In advancing age, the branches assume a drooping characteristic. The tree's pewter-colored rippled bark adds to the attractive aesthetic of this species. Typically, the leaves are long and are roundly and deeply lobed. The leaf width is approximately one half its length. Each leaf is matte green with an underneath pale green appearance; moreover, the leaf is covered with abundant soft fuzz, yielding an almost velvety feeling. When a fresh leaf is rubbed or broken, an
aroma An odor (American English) or odour (Commonwealth English; see spelling differences) is caused by one or more volatilized chemical compounds that are generally found in low concentrations that humans and animals can perceive via their sense ...
tic scent is exuded, evoking a forest
odor An odor (American English) or odour (Commonwealth English; see spelling differences) is caused by one or more volatilized chemical compounds that are generally found in low concentrations that humans and animals can perceive via their sense ...
. The wood is a dull brown approaching yellow. The
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s are medium to dark brown and range from in length. The caps have deep stippling and are found most often as singlets, but occasionally as doublets. The acorns ripen from October to November. Viable acorns germinate in their first winter, and none remain by mid-winter.


Taxonomy

Valley oak is of the
white oak The genus ''Quercus'' contains about 500 species, some of which are listed here. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the two subgenera '' ...
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
ary lineage, which is officially known as the
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''Lepidobalanus''. This subgenus comprises numerous oaks from California and elsewhere, which species share similar leaves, acorns, bark and
wood pulp Pulp is a lignocellulosic fibrous material prepared by chemically or mechanically separating cellulose fibers from wood, fiber crops, waste paper, or rags. Mixed with water and other chemical or plant-based additives, pulp is the major raw mate ...
. Early
settlers A settler is a person who has migrated to an area and established a permanent residence there, often to colonize the area. A settler who migrates to an area previously uninhabited or sparsely inhabited may be described as a pioneer. Settle ...
used a variety of common names for the valley oak including: white oak, bottom oak, swamp oak, water oak and mush oak. The
Spaniard Spaniards, or Spanish people, are a Romance ethnic group native to Spain. Within Spain, there are a number of national and regional ethnic identities that reflect the country's complex history, including a number of different languages, both ind ...
s, because the tree looked like the white oaks in Europe, called the tree "roble". The Concow tribe call the acorns lō-ē’ (
Konkow language The Konkow language, also known as Northwest Maidu (also ''Concow-Maidu'', or ' in the language itself) is a part of the Maiduan language group. It is spoken in California. It is severely endangered, with three remaining elders who learned to ...
).


Distribution and habitat

Valley oak tolerates cool wet winters and hot dry summers, but requires abundant water. It is most abundant in rich deep soils of valley floors below 600 meters (2000 feet) in elevation. Valley oak is found in dense
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
forests, open
foothill woodland Foothill woodland is a California vegetation type characterized by an overstory of broadleaf and coniferous trees, generally found in areas of higher elevations and more precipitation than grasslands and oak savannah, and at lower elevations and le ...
s and valley
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s. Commonly associated trees are
coast live oak ''Quercus agrifolia'', the California live oak, or coast live oak, is a highly variable, often evergreen oak tree, a type of live oak, native to the California Floristic Province. It may be shrubby, depending on age and growing location, but is ...
,
interior live oak ''Quercus wislizeni'', known by the common name interior live oak, is an evergreen oak, highly variable and often shrubby, found in many areas of California in the United States continuing south into northern Baja California in Mexico. It general ...
,
blue oak ''Quercus douglasii'', known as blue oak, is a species of oak endemic to (and found only in) California, common in the Coast Ranges and the foothills of the Sierra Nevada. It is California's most drought-tolerant deciduous oak, and is a dominan ...
, California black walnut,
California sycamore ''Platanus racemosa'' is a species of plane tree known by several common names, including California sycamore, western sycamore, California plane tree, and in North American Spanish aliso. ''Platanus racemosa'' is native to California and Baja ...
and
gray pine ''Pinus sabiniana'' (sometimes spelled ''P. sabineana''), with vernacular names including towani pine, foothill pine, gray pine, bull pine, and digger pine, is a pine endemic to California in the United States. Some sources discourage using the ...
. The valley oak is widely distributed in: the California Central Valley; many smaller valleys such as the
San Fernando Valley The San Fernando Valley, known locally as the Valley, is an urbanized valley in Los Angeles County, California. Located to the north of the Los Angeles Basin, it contains a large portion of the City of Los Angeles, as well as unincorporated ar ...
(original Spanish place-name from oak savannah),
Conejo Valley The Conejo Valley is a region spanning both southeastern Ventura County and northwestern Los Angeles County in Southern California, United States. It is located in the northwestern part of the Greater Los Angeles Area. Communities in the Con ...
, and
Santa Ynez Valley The Santa Ynez Valley is located in Santa Barbara County, California, between the Santa Ynez Mountains to the south and the San Rafael Mountains to the north. The Santa Ynez River flows through the valley from east to west. The Santa Ynez Valle ...
; the Inner Coast Ranges south of the Eel River; and the
Transverse Ranges The Transverse Ranges are a group of mountain ranges of southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa ...
from the
Tehachapi Mountains The Tehachapi Mountains (; Kawaiisu: ''Tihachipia'', meaning "hard climb") are a mountain range in the Transverse Ranges system of California in the Western United States. The range extends for approximately in southern Kern County and northw ...
to the
Simi Hills The Simi Hills are a low rocky mountain range of the Transverse Ranges in eastern Ventura County and western Los Angeles County, of southern California, United States. Geography The Simi Hills are aligned east-west and run for , and average aro ...
,
Santa Susana Mountains The Santa Susana Mountains are a transverse range of mountains in Southern California, north of the city of Los Angeles, in the United States. The range runs east-west, separating the San Fernando and Simi valleys on its south from the Santa ...
. It is also present on
Santa Cruz Island Santa Cruz Island (Spanish: ''Isla Santa Cruz'', Chumash: ''Limuw'') is located off the southwestern coast of Ventura, California, United States. It is the largest island in California and largest of the eight islands in the Channel Islands ...
and Catalina Island in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
. Some of the most picturesque stands are found in
Sonoma Valley Sonoma Valley is a valley located in southeastern Sonoma County, California, in the North Bay region of the San Francisco Bay Area. Known as the birthplace of the California wine industry, the valley is home to some of the earliest vineyards ...
, Round Valley in
Mendocino County Mendocino County (; ''Mendocino'', Spanish for "of Mendoza) is a county located on the North Coast of the U.S. state of California. As of the 2020 census, the population was 91,601. The county seat is Ukiah. Mendocino County consists whol ...
and the southern
Salinas Valley The Salinas Valley is one of the major valleys and most productive Agriculture, agricultural regions in California. It is located west of the San Joaquin Valley and south of San Francisco Bay and the Santa Clara Valley. The Salinas River (Califo ...
near the up-river reaches of the Salinas River.


Ecology

Like many oaks, valley oaks can tolerate wildfires. Although smaller individuals may be top-killed, most resprout from the root crown. A variety of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s and birds eat the acorns, including the
acorn woodpecker The acorn woodpecker (''Melanerpes formicivorus'') is a medium-sized woodpecker, long, with an average weight of . Taxonomy The acorn woodpecker was formally described in 1827 by the English naturalist William John Swainson under the binomia ...
,
California scrub jay The California scrub jay (''Aphelocoma californica'') is a species of scrub jay native to western North America. It ranges from southern British Columbia throughout California and western Nevada near Reno to west of the Sierra Nevada. The Cali ...
,
yellow-billed magpie The yellow-billed magpie ''(Pica nuttalli)'', also known as the California magpie, is a large bird in the crow family that is restricted to the U.S. state of California. It inhabits the Central Valley and the adjacent chaparral foothills and ...
, and
California ground squirrel The California ground squirrel (''Otospermophilus beecheyi''), also known as the Beechey ground squirrel, is a common and easily observed ground squirrel of the western United States and the Baja California Peninsula; it is common in Oregon and ...
. The acorns are also attacked by bruchid beetles, but can survive moderate levels of infestation. Globular
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants, fungi, or animals. Plant galls are abnormal outgrowths of plant tissues, similar to be ...
s up to several centimeters in diameter are frequently attached to twigs of mature specimens of valley oak. These house the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
l stage of small indigenous
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s '' Andricus quercuscalifornicus''. A related wasp species, '' A. kingi'', produces small galls shaped like Hershey's kisses on leaf surfaces. The valley oak is the only known food plant of '' Chionodes petalumensis'' caterpillars.


Uses

The acorns are sweet and edible; Native Americans including the
Southern Paiute people The Southern Paiute people are a tribe of Native Americans who have lived in the Colorado River basin of southern Nevada, northern Arizona, and southern Utah. Bands of Southern Paiute live in scattered locations throughout this territory and ha ...
roasted them and ground the edible portion into meal to make into bread and mush. Difficulties in acquiring valley oak wood as well as issues stemming from its drying such as cracking and warping have shifted its consumption from a general purpose lumber to a primarily niche product. Valley oak wood has a small, but significant market for use in cabinetry though, and is also suitable for hardwood flooring.
Tylose Tyloses are outgrowths/extragrouth on parenchyma cells of xylem vessels of secondary heartwood. When the plant is stressed by drought or infection, tyloses will fall from the sides of the cells and "dam" up the vascular tissue to prevent furth ...
s present in the pores of valley oak wood increase its impermeability to fluids allowing it to be used in the production of water-tight vessels. Such vessels include wine barrels where valley oak wood sees limited role in the composition of and where it has similar properties to other
white oaks The genus ''Quercus'' contains about 500 species, some of which are listed here. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the two subgenera '' ...
such as a reduced tannin load compared to the
red oaks The genus ''Quercus'' contains about 500 species, some of which are listed here. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the two subgenera ''C ...
and an open grain that allows for an increased transfer of oxygen.


Observational history

In 1792, the English explorer
George Vancouver Captain George Vancouver (22 June 1757 – 10 May 1798) was a British Royal Navy officer best known for his 1791–1795 expedition, which explored and charted North America's northwestern Pacific Coast regions, including the coasts of what a ...
noted on his expedition through the
Santa Clara Valley The Santa Clara Valley is a geologic trough in Northern California that extends 90 miles (145 km) south–southeast from San Francisco to Hollister. The longitudinal valley is bordered on the west by the Santa Cruz Mountains and on the ea ...
, after seeing an expanse of valley oaks: In the year 1861,
William Henry Brewer William Henry Brewer (September 14, 1828 – November 2, 1910) was an American botanist. He worked on the first California Geological Survey and was the first Chair of Agriculture at Yale University's Sheffield Scientific School. Biography Will ...
, the chief botanist for the first
California Geological Survey The California Geological Survey, previously known as the California Division of Mines and Geology, is the California state geologic agency. History Although it was not until 1880 that the California State Mining Bureau, predecessor to the Cal ...
wrote of the valley oaks that he saw in
Monterey County Monterey County ( ), officially the County of Monterey, is a county located on the Pacific coast in the U.S. state of California. As of the 2020 census, its population was 439,035. The county's largest city and county seat is Salinas. Montere ...
: The Hooker Oak of
Chico, California Chico ( ; Spanish for "little") is the most populous city in Butte County, California. Located in the Sacramento Valley region of Northern California, the city had a population of 101,475 in the 2020 census, reflecting an increase from 86,187 ...
, was once considered the largest-known valley oak. When it fell on May 1, 1977, it was nearly and in circumference at from the ground.


See also

*
California oak woodland California oak woodland is a plant community found throughout the California chaparral and woodlands ecoregion of California in the United States and northwestern Baja California in Mexico. Oak woodland is widespread at lower elevations in coastal ...
*
List of Quercus species The genus ''Quercus'' contains about 500 species, some of which are listed here. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the two subgenera '' ...


References


External links

* * {{Authority control
lobata Lobata is an order of Ctenophora in the class Tentaculata with smaller tentacles than other ctenophores, and distinctive flattened lobes extending outwards from their bodies. They grow up to about long. Anatomy The lobates have a pair of lo ...
Endemic flora of California Trees of the Southwestern United States Natural history of the California chaparral and woodlands Natural history of the California Coast Ranges Natural history of the Central Valley (California) Natural history of the Channel Islands of California Natural history of the San Francisco Bay Area Natural history of the Santa Monica Mountains Natural history of the Transverse Ranges Trees of Mediterranean climate Plants described in 1801 Garden plants of North America Drought-tolerant trees Ornamental trees