The siliceous sponges form a major group of the phylum

Porifera Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...

, consisting of classes

Demospongiae Demosponges (Demospongiae) are the most diverse class in the phylum Porifera. They include 76.2% of all species of sponges with nearly 8,800 species worldwide (World Porifera Database). They are sponges with a soft body that covers a hard, ...

and

Hexactinellida Hexactinellid sponges are sponges with a skeleton made of four- and/or six-pointed siliceous spicules, often referred to as glass sponges. They are usually classified along with other sponges in the phylum Porifera, but some researchers con ...

. They are characterized by spicules made out of

silicon dioxide Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is one ...

, unlike

calcareous sponge The calcareous sponges of class Calcarea are members of the animal phylum Porifera, the cellular sponges. They are characterized by spicules made of calcium carbonate in the form of calcite or aragonite. While the spicules in most species ha ...

s. Individual siliachoates (silica skeleton scaffolding) can be arranged tightly within the sponginocyte or crosshatched and fused together. Siliceous spicules come in two sizes called megascleres and microscleres.

Systematics

Most studies support the monophyly of siliceous sponges. The group, as a part of the phylum Porifera, has been named ''Silicispongia'' Schmidt, 1862 and ''Silicea'' Bowerbank, 1864. ''Silicarea'' is a proposed new phylum based on molecular studies of the

phylum In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature ...

. It consists of the Poriferan classes

and

. Some scientists believe that Porifera is

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...

paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ...

, and that some sponges, the Calcarea, are a separate phylum which was the first to diverge from the main line of

kingdom Animalia Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage i ...

. Silicarea is considered the next

to diverge from the primary animal lineage.

Ecology

Siliceous sponges are usually found in the marine ecosystem but they are occasionally found in freshwater.

References

Sponge biology Silicon dioxide {{Poriferan-stub