
Shrubland, scrubland, scrub, brush, or bush is a
plant community
A plant community is a collection or Association (ecology), association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The compo ...
characterized by
vegetation
Vegetation is an assemblage of plant
Plants are predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular re ...

dominated by
shrub
A shrub (often called a bush) is a small- to medium-sized perennial
A perennial plant or simply perennial is a plant
Plants are predominantly photosynthetic eukaryotes of the Kingdom (biology), kingdom Plantae. Historically, the p ...

s, often also including
grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family
In , family (from la, familia) is a of people related either by (by recognized birth) or (by marriage or other relationship). The purpose of families is to maintain ...

es,
herb
In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables
Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is app ...
s, and
geophyte
A storage organ is a part of a plant
Plants are mainly multicellular organisms, predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical ene ...
s. Shrubland may either occur naturally or be the result of human activity. It may be the mature
vegetation type
Vegetation classification is the process of classifying and mapping the vegetation over an area of the earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management. ...
in a particular region and remain stable over time, or a transitional community that occurs temporarily as the result of a disturbance, such as fire. A stable state may be maintained by regular natural disturbance such as fire or
browsing
Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor
A metaphor is a figure of speech that, for rhetorical effect, directly refers ...
. Shrubland may be unsuitable for human habitation because of the danger of fire. The term was coined in 1903.
Shrubland species generally show a wide range of
adaptations
In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process that fits organisms to their environment, enhancing their Fitness (biology), evolutionary fitness. Secondly, it is a state reached by the popula ...
to fire, such as heavy seed production,
lignotubers, and fire-induced germination.
Botanical structural form
In
botany
Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...

and
ecology
Ecology (from el, οἶκος, "house" and el, -λογία, label=none, "study of") is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms
In biol ...
a shrub is defined as a much-branched
woody plant
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to Herbaceous plant, herbaceous plants that die back to t ...
less than 8 m high and usually with many
. Tall shrubs are mostly 2–8 m high, small shrubs 1–2 m high and
subshrub
A subshrub (Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally spoken in the area around Rome, known as Latium. Through the power of the Roman Republic, it ...

s less than 1 m high.
A descriptive system widely adopted in
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

to describe different types of vegetation is based on
structural characteristics based on
plant life-formPlant life-form schemes constitute a way of classifying plants alternatively to the ordinary species-genus-family scientific classification
In biology
Biology is the natural science that studies life and living organisms, including thei ...
, plus the height and foliage cover of the tallest stratum or
dominant
Domination or dominant may refer to:
Society
* World domination, which is mainly a conspiracy theory
* Colonialism in which one group (usually a nation) invades another region for material gain or to eliminate competition
* Chauvinism in which a p ...
species
In biology, a species is the basic unit of biological classification, classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individu ...

.

For shrubs 2–8 m high the following structural forms result:
* dense foliage cover (70–100%) — closed-scrub
* mid-dense foliage cover (30–70%) — open-scrub
* very sparse foliage cover (<10%) — tall open shrubland
For shrubs <2 m high the following structural forms result:
* dense foliage cover (70–100%) — closed-
heath
A heath () is a shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, Herbaceous plant, herbs, and geophytes. Shrubland m ...
* mid-dense foliage cover (30–70%) — open-heath
* sparse foliage cover (10–30%) — low shrubland
* very sparse foliage cover (<10%) — low open shrubland
Biome plant group

Similarly, shrubland is a category used to describe a type of
biome
A biome is a collection of plants
Plants are predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respi ...
plant group. In this context, shrublands are dense thickets of evergreen
sclerophyll
Sclerophyll is a type of vegetation that has hard leaves
A leaf (plural leaves) is the principal lateral appendage of the vascular plant
Vascular plants (from Latin ''vasculum'': duct), also known as Tracheophyta (the tracheophytes , ...
shrubs and small trees,
called:
*
Chaparral
Chaparral ( ) is a shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, Herbaceous plant, herbs, and geophytes. Shrublan ...

in
California
California is a state
State may refer to:
Arts, entertainment, and media Literature
* ''State Magazine'', a monthly magazine published by the U.S. Department of State
* The State (newspaper), ''The State'' (newspaper), a daily newspaper i ...

*
Matorral
image:Matorral Octubre.JPG, 300px, Springtime in Chilean matorral a few kilometers north of Santiago along the Pan-American Highway
Matorral is a Spanish language, Spanish word, along with ''tomillares'', for shrubland, thicket or bushes. It is us ...
in
Chile
Chile, officially the Republic of Chile, is a country in the western part of South America
South America is a entirely in the and mostly in the , with a relatively small portion in the . It can also be described as the southern ...

,
Mexico
Mexico, officially the United Mexican States, is a country
A country is a distinct territorial body or political entity
A polity is an identifiable political entity—any group of people who have a collective identity, who are organi ...

, and
Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto =
, national_anthem =
, image_map =
, map_caption =
, image_map2 ...

*
Maquis in
France
France (), officially the French Republic (french: link=no, République française), is a transcontinental country
This is a list of countries located on more than one continent
A continent is one of several large landmasses ...

and elsewhere around the
Mediterranean
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western Europe, Western and Southern Europe and Anatolia, on the south by North Africa ...

*
in
Italy
Italy ( it, Italia ), officially the Italian Republic ( it, Repubblica Italiana, links=no ), is a country consisting of a peninsula delimited by the Alps
The Alps ; german: Alpen ; it, Alpi ; rm, Alps; sl, Alpe ) are the highest ...

*
Fynbos
Fynbos (; meaning fine plants) is a small belt of natural shrubland or Heath (habitat), heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. This area is predominantly coastal and mountainous, with a M ...

in
South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. With over 60 million people, it is the world's 23rd-most populous nation and covers an area of . South Africa has three capital citie ...

*
Cumberland Plain Woodland
The Cumberland Plain Woodland is one of six main indigenous woodland communities of Sydney
Sydney ( ; Dharug language, Dharug: ) is the List of Australian capital cities, capital city of the state of New South Wales, and the List of cities ...
in
Sydney
Sydney ( ; Dharug
The Darug or Dharug people are an Aboriginal Australian people, who share strong ties of kinship and, in Colonial Australia, pre-colonial times, survived as skilled hunters in family groups or clans, scattered througho ...

,
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

*
Kwongan
Kwongan is plant community found in south-western Western Australia
Western Australia (abbreviated as WA) is a States and territories of Australia, state occupying the western percent of the land area of Australia excluding external territori ...
in Southwest
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

*
Cedar scrub in
Texas Hill Country
The Texas Hill Country is a geographic region of Central and South Texas, forming the southeast part of the Edwards Plateau. Given its location, climate, terrain, and vegetation, the Hill Country can be considered the border between the Amer ...

In some places shrubland is the mature vegetation type, and in other places the result of degradation of former
forest
A forest is an area of land dominated by tree
In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, supporting branches and leaves in most species. In some usages, the definition of a ...

or
woodland
A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum
A ''plurale tantum'' (Latin for "plural only"; ) is a noun that appears only in the plural
The plu ...

by
logging
Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidder, skidding, on-site processing, and loading of trees or trunk (botany), logs onto logging truck, trucks or flatcar#Skeleton car, s ...

or
overgrazing
Overgrazing occurs when plants are exposed to intensive grazing
In agriculture
Agriculture is the science, art and practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentism, sedentary hum ...

, or disturbance by major fires.
A number of
World Wildlife Fund
In its most general sense, the term "world" refers to the totality of entities, to the whole of reality or to everything that is. The nature of the world has been conceptualized differently in different fields. Some conceptions see the worl ...
biomes are characterized as shrublands, including:
[
;Desert scrublands
]
''Xeric'' or ''desert scrublands'' occur in the world's deserts and xeric shrublands
Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this ha ...
ecoregions, or in areas of fast-draining sandy soils in more humid regions. These scrublands are characterized by plants with adaptations to the dry climate, which include small leaves to limit water loss, thorns to protect them from grazing animals, succulent leaves or stems, storage organ
A storage organ is a part of a plant
Plants are predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respi ...
s to store water, and long taproots to reach groundwater.
;Mediterranean scrublands
''Mediterranean scrublands'' occur naturally in the Mediterranean forests, woodlands, and scrub
Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
biome
A biome is a collection of plants
Plants are predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy that, through cellular respi ...
s, located in the five Mediterranean climate
A Mediterranean climate or dry summer climate is characterized by dry summers and mild, wet winters. The climate
Climate is the long-term pattern of weather
Weather is the state of the atmosphere, describing for example the degre ...
regions of the world. Scrublands are most common near the seacoast, and have often adapted to the wind and salt air of the ocean. Low, soft-leaved scrublands around the Mediterranean Basin
In biogeography, the Mediterranean Basin (also known as the Mediterranean region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by ...
are known as ''garrigue
.
Garrigue or phrygana (n.pl., Greek Φρύγανα) is a type of low, scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome.
It is found on limestone
Limestone is a common type of carbonate r ...
'' in France
France (), officially the French Republic (french: link=no, République française), is a transcontinental country
This is a list of countries located on more than one continent
A continent is one of several large landmasses ...

, ''phrygana'' in Greece
Greece ( el, Ελλάδα, Elláda, ), officially the Hellenic Republic, is a country located in Southeastern Europe
Southeast Europe or Southeastern Europe () is a geographical subregion
A subregion is a part of a larger region
In geogr ...

, ''tomillares'' in Spain
, image_flag = Bandera de España.svg
, image_coat = Escudo de España (mazonado).svg
, national_motto =
, national_anthem =
, image_map =
, map_caption =
, image_map2 ...

, and ''batha'' in Israel
Israel (; he, יִשְׂרָאֵל, translit=Yīsrāʾēl; ar, إِسْرَائِيل, translit=ʾIsrāʾīl), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a ...

. Northern coastal scrub
Northern coastal scrub is a scrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, Herbaceous plant, herbs, and geophytes. Shr ...
and coastal sage scrub
Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community
A plant community is a collection or Association (ecology), association of plant spec ...
occur along the California
California is a state
State may refer to:
Arts, entertainment, and media Literature
* ''State Magazine'', a monthly magazine published by the U.S. Department of State
* The State (newspaper), ''The State'' (newspaper), a daily newspaper i ...

coast, strandveld in the Western Cape
The Western Cape ( af, Wes-Kaap; xh, iNtshona-Koloni) is a province
A province is almost always an administrative division
Administrative division, administrative unitArticle 3(1). , country subdivision, administrative region, subnation ...
of South Africa
South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. With over 60 million people, it is the world's 23rd-most populous nation and covers an area of . South Africa has three capital citie ...

, coastal matorral in central Chile
Central Chile (''Zona central'') is one of the five natural regions into which CORFO divided continental Chile in 1950. It is home to a majority of the Chile
Chile (, ; ), officially the Republic of Chile (), is a country in western Sou ...
, and sand-heath and kwongan
Kwongan is plant community found in south-western Western Australia
Western Australia (abbreviated as WA) is a States and territories of Australia, state occupying the western percent of the land area of Australia excluding external territori ...
in Southwest Australia
Southwest Australia is a ecoregion, biogeographic region in Western Australia. It includes the Mediterranean climate, Mediterranean-climate area of southwestern Australia, which is home to a diverse and distinctive flora and fauna.
The region is ...
.
;Interior scrublands
''Interior scrublands'' occur naturally in semi-arid areas where soils are nutrient-poor, such as on the matas of Portugal
Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country
A country is a distinct territorial body or political entity
A polity is an identifiable political entity—any group of people who ...

which are underlain by Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period
A geological period is one of the several subdivisions of geologic time enabling cross-referencing of rocks and geologic events from place to place.
These peri ...
and Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician
The Ordovician ( ) is a geologic period
A geological period is one of the several subdivisions of geologic time enabling cross-refere ...
schists
Schist ( ) is a medium-grained metamorphic rock
, a type of metamorphic rock
Metamorphic rocks arise from the transformation of existing rock (geology), rock to new types of rock, in a process called metamorphism
upright=1.35, Schematic r ...
. Florida scrub
Florida sand pine scrub is an endangered subtropical forest ecoregion found throughout Florida in the United States. It is found on coastal and inland sand ridges and is characterized by an evergreen Xerophyte, xeromorphic plant community dominated ...
is another example of interior scrublands.
;Dwarf shrubs
Some vegetation types are formed of '' dwarf-shrubs'': low-growing or creeping shrubs. These include the maquis and garrigue
.
Garrigue or phrygana (n.pl., Greek Φρύγανα) is a type of low, scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome.
It is found on limestone
Limestone is a common type of carbonate r ...
s of Mediterranean climates, and the acid-loving dwarf shrubs of heathland
A heath () is a shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community
A plant community is a collection or Association (ecology), association of plant species within a designated geographical unit, which forms a relativ ...
and moorland
Moorland or moor is a type of habitat
In ecology
Ecology (from el, οἶκος, "house" and el, -λογία, label=none, "study of") is the study of the relationships between living organisms, including humans, and their physi ...
.
See also
* Fynbos
Fynbos (; meaning fine plants) is a small belt of natural shrubland or Heath (habitat), heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. This area is predominantly coastal and mountainous, with a M ...

* Maquis
* Prostrate shrub
A prostrate shrub is a woody plant
A woody plant is a plant
Plants are predominantly photosynthetic
Photosynthesis is a process used by plants and other organisms to Energy transformation, convert light energy into chemical energy t ...
* Semi-desert
A semi-arid climate, semi-desert climate, or steppe climate is the climate
Climate is the long-term pattern of weather
Weather is the state of the atmosphere, describing for example the degree to which it is hot or cold, wet or dry, c ...
* Shrub-steppe
* Shrub swamp
Shrub swamps — also called scrub swamps or buttonbush swamps — are a type of freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water
Water (chemical formula H2O) is an , transparent, tasteless, o ...

* Moorland
Moorland or moor is a type of habitat
In ecology
Ecology (from el, οἶκος, "house" and el, -λογία, label=none, "study of") is the study of the relationships between living organisms, including humans, and their physi ...
Notes and references
{{Vegetation