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The scalloped hammerhead (''Sphyrna lewini'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
hammerhead shark The hammerhead sharks are a group of sharks that form the family Sphyrnidae, so named for the unusual and distinctive structure of their heads, which are flattened and laterally extended into a "hammer" shape called a cephalofoil. Most hammerhe ...
in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Sphyrnidae. It was originally known as ''Zygaena lewini''. The
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''sphyrna'' translates into "hammer" in English, referring to the shape of this shark's head, which is its most distinguishing characteristic. The shark's eyes and nostrils are at the tips of the extensions. It is a fairly large hammerhead, but is still smaller than both the
great Great may refer to: Descriptions or measurements * Great, a relative measurement in physical space, see Size * Greatness, being divine, majestic, superior, majestic, or transcendent People * List of people known as "the Great" *Artel Great (born ...
and smooth hammerheads. This shark is also known as the bronze, kinky-headed, or southern hammerhead. It primarily lives in warm, temperate, and tropical coastal waters all around the globe between
latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north ...
s 46°N and 36°S, down to a depth of . It is the most common of all hammerheads.


Taxonomy

The scalloped hammerhead was first named ''Zygaena lewini'' and then renamed ''Sphyrna lewini'' by Edward Griffith and Hamilton Smith in 1834. It has also been named ''Cestracion leeuwenii'' by Day in 1865, ''Zygaena erythraea'' by Klunzinger in 1871, ''Cestracion oceanica'' by Garman in 1913, and ''Sphyrna diplana'' by Springer in 1941. ''Sphyrna'' comes from the Greek and translates to hammer. It is a sister species to '' Sphyrna gilberti'', differing by the number of vertebrae. Though once considered a distinct species, McEachran and Serret synonymized ''Sphyrna couardi'' with ''Sphyrna lewini'' in 1986.


Description

Typically, males measure and weigh about when they attain sexual maturity, whereas the larger females measure and weigh about at sexual maturity. The maximum length of the scalloped hammerhead is and the maximum weight is , per
FishBase FishBase is a global species database of fish species (specifically finfish). It is the largest and most extensively accessed online database on adult finfish on the web.
. A female caught off of
Miami Miami ( ), officially the City of Miami, known as "the 305", "The Magic City", and "Gateway to the Americas", is a coastal metropolis and the county seat of Miami-Dade County in South Florida, United States. With a population of 442,241 at ...
was found to have measured and reportedly weighed , though was in a
gravid In biology and human medicine, gravidity and parity are the number of times a woman is or has been pregnant (gravidity) and carried the pregnancies to a viable gestational age (parity). These terms are usually coupled, sometimes with additional t ...
state then.Castro, José I. (2011) ''The Sharks of North America''. Oxford University Press. These sharks have a very high metabolic rate, which governs behavior in acquiring food. They occupy tertiary trophic levels. The scalloped hammerhead shark, like many other species, uses the shore as a breeding ground. Due to high metabolic rates, young scalloped hammerhead sharks need a lot of food, or they will starve. ''S. lewini'' is also noted for its large and complex brain, with high levels of cognitive capabilities, social intelligence, sensorimotor functions, intricate migrations, complex habitat relationships, and athletic capture of prey. Their social intelligence is especially important for their aggregative behavior, allowing the species to reproduce with the fittest members and proliferate the species easier. Sphyrna lewini upper teeth.jpg, Upper teeth Sphyrna lewini lower teeth.jpg, Lower teeth Sphyrna lewini juvenile.jpg, Juvenile Sphyrna lewini young.jpg, Juvenile


Distribution and habitat

The scalloped hammerhead is a coastal
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
species; it occurs over continental and insular shelves and in nearby deeper water. It is found in warm temperate and
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
waters, worldwide from 46°N to 36°S. It can be found down to depths over , but is most often found above . During the day, they are more often found close to shore, and at night, they hunt further offshore. Adults are found alone, in pairs, or in small
schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...
, while young sharks occur in larger schools. Juveniles and pups thrive in shallow coastal waters, such as bays and mangroves, which provide shelter from predators and waters high in nutrients from deposited sediments. Research carried out by the nongovernmental organisation
Misión Tiburón Misión Tiburón is a non profit organisation established in 2009, based on the Pacific coast of Costa Rica. Founded by marine biologists and conservationists Ilena Zanella and Andres Lopez, the organisation focuses on marine research concerning ...
, using conventional and acoustic shark tagging methods, found that adult scalloped hammerheads migrate from the pelagic waters surrounding Cocos Island to the mangroves in the tropical fjord of Golfo Dulce - a tropical fjord on the Pacific Coast of Costa Rica. Here, female sharks give birth to live young; juveniles remain in the shallow root system of the mangroves for around three years. After this time they leave Golfo Dulce and migrate back to Cocos Island, to feed in pelagic waters.


Behavior


Schooling

These sharks are often seen during the night, day, and morning in big schools, sometimes numbering hundreds, most likely because large groups can obtain food easier than singles or small groups, especially larger and trickier prey, as commonly seen. The younger the sharks, the closer to the surface they tend to be, while the adults are found much deeper in the ocean. They are not considered dangerous and are normally not aggressive towards humans.


Sexual dimorphism

The female scalloped hammerheads migrate offshore at a smaller size than males because the larger classes of the hammerhead, such as those from 100 to 140 cm long, travel deeper down. Males and females differ in that males are observed to stay deeper than female sharks in general. Sexual maturity generally occurs once the scalloped hammerhead attains 240 cm in total or longer. Physically, the mature females have considerably wider uteri than their maturing counterparts. A lack of mating scars has been found on mature females. Unlike females, males reach sexual maturity at a much smaller size. The male-to-female ratio of the scalloped hammerhead is 1:1.29. Females probably are capable of giving birth annually. usually in the summer.


Navigating behavior

Scalloped hammerhead sharks have a homing behavior to navigate in the ocean. They move in the night and use the environment as a map, similar to a human reading a topographical map. By experimentation in tagging these sharks, one could test for any guidance in a shark's movement. These sharks use a point-to-point type of school swimming, and do not favor going too deep, where temperature changes hitchhike with current speed and directional change. The scalloped hammerhead uses deep water to survive as safety and feeding. Although they have high metabolic rates, they have a tendency to be sedentary and allow currents to carry them as they swim. As a result, this causes scalloped hammerheads to be selective where they swim and the depth at which they tend to stay. They also make use of earths magnetic field.


Predation

The scalloped hammerhead has several advantages to capture its prey. The shape of its head allows it to bury into the seafloor and pin stingrays down. The wide head and special sensory cells allow the scalloped hammerhead to successfully detect fish.


Reproduction

The gestation period is reported to be around 12 months. Scalloped hammerheads give live birth. Compared to other species, the scalloped hammerhead produces large litters (12-41 pups), and this is most likely due to high infant mortality. Like most sharks, parental care is not seen. Nursery grounds for this species are predictable and repeated over the years, and they are faithful to their natal sites. Their natal sites still cause high infant mortality; a lack of resources prevents all the young from surviving. As a result, only the fittest grow to maturity. Also, should a population get depleted, it recovers through reproduction and not immigration. This species does not seem to attack each other even in periods of starvation. In addition, scalloped hammerheads have migratory behaviors. As a result, deprivation results from migration and young growth. While the Taiwan scalloped hammerhead seems to have an earlier maturity rate, it is still reported to be slow to mature.


Diet

Scalloped hammerheads are generalists and opportunistic predators, eating whatever is available and abundant in their area. There is no difference in what male and female sharks eat, but larger sharks are noted to eat larger prey than smaller, juvenile hammerheads. Juveniles typically feed on inshore fish, while adults feed on inshore fish as well as larger organisms that live in deeper waters. Specifically, this shark feeds primarily on fish such as
sardine "Sardine" and "pilchard" are common names for various species of small, oily forage fish in the herring family Clupeidae. The term "sardine" was first used in English during the early 15th century, a folk etymology says it comes from the It ...
s,
mackerel Mackerel is a common name applied to a number of different species of pelagic fish, mostly from the family Scombridae. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. ...
, and
herring Herring are forage fish, mostly belonging to the family of Clupeidae. Herring often move in large schools around fishing banks and near the coast, found particularly in shallow, temperate waters of the North Pacific and North Atlantic Ocean ...
, and occasionally they feed on cephalopods such as
squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fittin ...
and
octopus An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, ...
. Larger specimens may also feed on smaller species of shark such as the
blacktip reef shark The blacktip reef shark (''Carcharhinus melanopterus'') is a species of requiem shark, in the family Carcharhinidae, which can be easily identified by the prominent black tips on its fins (especially on the first dorsal fin and its caudal fin). ...
, ''Carcharhinus melanopterus''.


Conservation

As of 2021, the scalloped hammerhead has been categorized as Critically Endangered by the IUCN Red List. The IUCN cites overfishing as the main cause for the drop in population numbers. Scalloped hammerheads are over harvested for their large sizes and fins that have 'high fin needle content;' their fins are sold for US $100–120 per kilogram. Many conservation efforts have been taken to protect this species, such as using genetic information from fins purchased from a Hong Kong market to pinpoint where scalloped hammerheads are being caught and track their exploitation levels . Another important effort has been protecting ''Sphyrna lewini'' nurseries. Scalloped hammerheads return to their birthplace to have their own pups; the pups will then spend their first years of life in these coastal nursery areas where they are safe from predators and have less competition for feeding before they venture into the open ocean. Fishing bans have been placed on some of these areas, such as in the Western Yucatan Peninsula, during breeding seasons to protect the young and juvenile scalloped hammerheads. In parts of the Atlantic Ocean, their populations had declined by over 95% in the past 30 years. Among the reasons for this drop are overfishing and the rise in demand for shark fins. Researchers attribute this growth in demand to the increase in shark fins as an expensive delicacy (such as in
shark fin soup Shark fin soup is a traditional soup or stewed dish served in parts of China, Taiwan, and Southeast Asia. The shark fins provide texture, while the taste comes from the other soup ingredients. It is commonly served at special occasions such as ...
) and in 2008 called for a ban on shark finning, a practice in which the shark's fins are cut off and the rest of the animal is thrown back in the water to die. Hammerheads are among the most commonly caught sharks for finning. "This species tendency to aggregate in large groups making capture in large numbers on long lines, bottom nets and trawls even easier." Hammerhead sharks are overfished all around the world for their fins and liver oil. an estimated 1.3 to 2.7 million fins are collected each year from smooth and scalloped hammerhead sharks for the shark-fin trade. According to a January 2021 study in ''
Nature Nature, in the broadest sense, is the physical world or universe. "Nature" can refer to the phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. Although humans are ...
'' which studied 31 species of sharks and rays, the number of these species found in open oceans had dropped by 71 per cent in around 50 years. The scalloped hammerhead was included in the study.


See also

*
List of sharks Shark is the naming term of all members of Selachimorpha suborder in the subclass Elasmobranchii, in the class Chondrichthyes. The Elasmobranchii also include rays and skates; the Chondrichthyes also include Chimaeras. The first shark-like chond ...


References

*


External links

* https://oceana.org/marine-life/sharks-rays/smooth-hammerhead-shark * * * ARKive �
images and movies of the scalloped hammerhead (''Sphyrna lewini'')



Species Description of ''Sphyrna lewini'' at www.shark-references.com
* {{Taxonbar, from=Q873701
scalloped hammerhead The scalloped hammerhead (''Sphyrna lewini'') is a species of hammerhead shark in the family Sphyrnidae. It was originally known as ''Zygaena lewini''. The Greek word ''sphyrna'' translates into "hammer" in English, referring to the shape of thi ...
Pantropical fish
scalloped hammerhead The scalloped hammerhead (''Sphyrna lewini'') is a species of hammerhead shark in the family Sphyrnidae. It was originally known as ''Zygaena lewini''. The Greek word ''sphyrna'' translates into "hammer" in English, referring to the shape of thi ...
scalloped hammerhead The scalloped hammerhead (''Sphyrna lewini'') is a species of hammerhead shark in the family Sphyrnidae. It was originally known as ''Zygaena lewini''. The Greek word ''sphyrna'' translates into "hammer" in English, referring to the shape of thi ...
Taxa named by Charles Hamilton Smith Species endangered by use as food Fish of the Dominican Republic