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The sauger (''Sander canadensis'') is a
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...
perciform fish of the family
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more ...
that resembles its close relative, the walleye. The species is a member of the largest vertebrate order, the Perciformes.Jaeger, Matthew. 2004. Montana's Fish Species of Special Concern: Sauger. Montana Cooperative Fisheries Research Unit. http://www.fisheriessociety.org/AFSmontana/SSCpages/Sauger%20Status.htm. (Accessed May 2011). It is the most migratory percid species in North America.Kuhn et al. 2008. Habitat Use and Movement Patterns by Adult Saugers from fall to Summer in an Unimpounded Small-River System. ''North American Journal of Fisheries Management''. 28: 360-367. American Fisheries Society. 2008. Saugers have two dorsal fins; the first is spiny and the posterior dorsal fin is soft-rayed. Their paired fins are in the thoracic position and their caudal fin is truncated, which means squared off at the corners, a characteristic of the family Percidae. Another physical characteristic of saugers is their ctenoid scales, which are common in advanced fishes. Saugers have a fusiform body structure, and as a result are well adapted predatory fishes and are capable of swimming into fast currents with minimal drag on their bodies. They may be distinguished from walleyes by the distinctly spotted
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
, by the lack of a white splotch on the
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
, by the rough skin over their gills, and by their generally more brassy color, or darker (almost black) color in some regions. The typical sauger is in weight.


Distribution

Saugers are widely distributed; their historical range consisted of the eastern U.S west of the
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. The ...
, mostly southern, central, and western U.S., and north into southern Canada. Sauger distribution and range has decreased from historical ranges because of degraded and fragmented habitat conditions. Sauger distribution within their home range varies by time of year because they are migratory. Saugers are more typical in rivers, whereas walleyes are more common in lakes and reservoirs. In many parts of their range, saugers are sympatric with walleyes.
Hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
between saugers and walleyes is not unknown; the hybrids, referred to as saugeyes, exhibit traits of both species. Being intermediate in appearance between the two species, saugeyes are sometimes difficult to differentiate, but they generally carry the dark blotches characteristic of the sauger. Saugers, however, are usually smaller and better tolerate waters of higher turbidity than walleyes. Saugers require warmer summer water temperatures of 20-28°C. Their need for warm water is thought to affect the northern and western boundaries of their range.


Reproduction and lifecycle

Saugers generally move upstream to spawn during March–May, depending on where they are. They move downstream to their home locations from April–July after their spawning period is over. Saugers have been known to travel between 10 and 600 km from their home to spawning locations downstream. Habitats at spawning sites are less complex and diverse than home locations. Females prefer rocky substrate and pools to deposit their eggs. As females increase in length, egg quality and fecundity increase, but egg production is thought to decline after age 6.Graeb, Brian D.S., Mark A. Kaemingk, David W. Willis. 2007. Early Life History of Sauger in Missouri River Reservoirs. Department of Wildlife and Fisheries Sciences. Brookings, SD. Sexual maturity is reached between 2 and 5 years old. Other measures of sexual maturity are related to size. A sauger is considered to be an adult when it reaches 250–300 mm. Upon hatching, larval saugers drift downstream before developing feeding tendencies and horizontal maneuverability. Juvenile saugers tend to develop in diversion canals and backwaters until autumn, when they migrate upstream to their winter habitat. Residing in diversion canals is a large cause of mortality for juvenile saugers.


Diet

Saugers feed on a variety of invertebrates and small fishes depending on the time of year and size of the sauger. Channel catfish ''Ictalurus punctatus'' and
Freshwater drum The freshwater drum, ''Aplodinotus grunniens'', is a fish endemic to North and Central America. It is the only species in the genus ''Aplodinotus'', and is a member of the family Sciaenidae. It is the only North American member of the group that ...
''Aplodinotus grunniens'' are a midsize (300–379 mm) and large (>379 mm) saugers' main food source during spring. The diet of a small sauger (200–299 mm) is slightly different from a larger sauger's. Smaller saugers generally feed on benthic invertebrates, mayfly larvae, and catfish during spring and summer. Midsized and large saugers feed mainly on fish from spring to autumn, but their diets alter during summer. Midsized and large saugers feed predominantly on mayfly larvae but only during summer. Freshwater drum ''Aplodinotus grunniens'' and gizzard shad ''Dorosoma cepedianum'' are predominant food sources for saugers of all sizes during autumn. Fish accounted for over 99% of a sauger's diet during autumn. Saugers also prey on shiners during spring and summer, but they do not account for a significant part of their diet. Shiners are absent from a sauger's diet in autumn probably due to their availability.Wickstrom, Gerald A. 2006. Seasonal Distribution, Movement, and Food Habits of Walleye and Sauger in Lewis and Clark Lake. South Dakota Department of Game, Fish & Parks. Chamberlain, South Dakota.


Habitat

Saugers are more likely to be found in large rivers with deep pools (depths greater than 0.6 m). They encounter a variety of habitats because of their migratory tendencies. They are usually found in natural rivers because they have more abundant pools and their flows have not been altered by dams or diversions. They are still common in impounded river systems. Diversions and dams affect habitat and spawning areas of saugers. They are usually found in areas with high turbidity, low channel slope, low stream velocity, and deep water. Saugers tend to select pools with sand and silt substrates, and habitat features that provide cover from the river current. They tend to avoid runs and riffles. They are most commonly found in pools that are at least 1.5 m deep. They can also be found in shallower pools, but in lesser numbers. No differences in habitat preference between males and females has been observed. The number of saugers observed increases with mean summer water temperature, maximum water depth, and alkalinity.


Taxonomy

Alongside the walleye, the sauger forms the North American
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
within the genus '' Sander'', and the sauger is thought to have first speciated into its modern form about 7.3 million years ago. The sauger was first formally described as ''Lucioperca canadensis'' in 1834 by British naturalists and explorers Edward Griffith (1790-1858) and Charles Hamilton Smith (1776-1859) based on the work of
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
naturalist and anatomist
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in na ...
(1769-1832) which Griffith translated from the
French language French ( or ) is a Romance language of the Indo-European family. It descended from the Vulgar Latin of the Roman Empire, as did all Romance languages. French evolved from Gallo-Romance, the Latin spoken in Gaul, and more specifically in N ...
.


Conservation

Saugers face many conservation issues because of migratory barriers,
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
, entrainment in
irrigation Irrigation (also referred to as watering) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has been devel ...
canals, and overexploitation. Dams and diversion canals prevent spawning in upstream habitats. Altering flows in rivers affect turbidity, formation of pools, and temperature, all of which are important for the timing and success of spawning saugers. Long migrations are the main reason why saugers struggle in dammed or diverted river systems. Low water levels in periods of drought are the most detrimental to sauger populations because it strands eggs during spawning and prevents larval saugers from reaching their downstream locations. High death rates that occur during spawning are related to degraded and fragmented river systems. Mortality rates in autumn are related to exploitation by fishermen. Sauger are critically imperiled in the state of New York, with a small population still remaining in
Lake Champlain , native_name_lang = , image = Champlainmap.svg , caption = Lake Champlain-River Richelieu watershed , image_bathymetry = , caption_bathymetry = , location = New York/ Vermont in the United States; and Quebec in Canada , coords = , type ...
. Sauger were extirpated from
Lake Erie Lake Erie ( "eerie") is the fourth largest lake by surface area of the five Great Lakes in North America and the eleventh-largest globally. It is the southernmost, shallowest, and smallest by volume of the Great Lakes and therefore also ha ...
at some point after a population crash in the 1950s, as well as from
Lake Ontario Lake Ontario is one of the five Great Lakes of North America. It is bounded on the north, west, and southwest by the Canadian province of Ontario, and on the south and east by the U.S. state of New York. The Canada–United States border sp ...
. Populations in the Allegheny River are prevented from reaching New York by the Kinzua Dam, which has led to the initiation of a sauger stocking program in the upper Allegheny River. As of 2022, fishing for sauger in New York is totally prohibited statewide.


References

*


External links


Fishbase information on Sauger
*Kansas Department of Wildlife and Park

*Montana Fish, Wildlife and Park

*Lake-Link.co

*Sauger Fishing in the Tennessee Valle
Saugeye
{{Taxonbar, from=Q1725670 Sander (fish) Freshwater fish of the United States Fish of the Eastern United States Fish of the Western United States Fish of the Great Lakes Fish described in 1834 Taxa named by Edward Griffith (zoologist)