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The rostellum is a projecting part of the
column A column or pillar in architecture and structural engineering is a structural element that transmits, through compression, the weight of the structure above to other structural elements below. In other words, a column is a compression member. ...
in
Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
flowers, and separates the male androecium from the female
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils' ...
, commonly preventing self-fertilisation. In many orchids, such as ''
Orchis mascula ''Orchis mascula'', the early-purple orchid, early spring orchis, is a species of flowering plant in the orchid family, Orchidaceae. Description ''Orchis mascula'' is a perennial herbaceous plant with stems up to high, green at the base a ...
'', the pollinia or pollen masses, are connected by stipes down to adhesive discs attached to the rostellum which forms cups keeping the discs or balls sticky. In '' Catasetum'' flowers the rostellum projects forward at each side as an "antenna", and the pollen masses are connected by a bent stalk or
pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...
to a sticky disc kept moist at the back of the flower. When an insect touches an "antenna", this releases the bent pedicel which springs straight and fires the pollinium, sticky disc first, at the insect.
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 â€“ 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended f ...
described in ''
Fertilisation of Orchids ''Fertilisation of Orchids'' is a book by English naturalist Charles Darwin published on 15 May 1862 under the full explanatory title ''On the Various Contrivances by Which British and Foreign Orchids Are Fertilised by Insects, and On the Good ...
'' how he "touched the antennæ of C. callosum whilst holding the flower at about a yard's distance from the window, and the pollinium hit the pane of glass, and adhered to the smooth vertical surface by its adhesive disc."


References

* *{{Citation , last =Darwin , first = Charles , year =1862 , title =On the various contrivances by which British and foreign orchids are fertilised by insects, and on the good effects of intercrossing , location = London , publisher = John Murray , url =http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F800&pageseq=1 , accessdate =2009-02-07 , postscript = Orchid morphology