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''Dacrydium cupressinum'', commonly known as rimu, is a large evergreen
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ...
tree
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the forests of New Zealand. It is a member of the southern
conifer Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...
group, the
podocarp Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pr ...
s. The Māori name ''rimu'' comes from the Polynesian term ''limu'' which the tree's foliage were reminded of, ultimately from
Proto-Austronesian Proto-Austronesian (commonly abbreviated as PAN or PAn) is a proto-language. It is the reconstructed ancestor of the Austronesian languages, one of the world's major language families. Proto-Austronesian is assumed to have begun to diversify ...
*''limut'' meaning "moss". The former name "red pine" has fallen out of common use.


Distribution

Rimu grows throughout New Zealand, in the
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-largest ...
,
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman S ...
and
Stewart Island/Rakiura Stewart Island ( mi, Rakiura, ' glowing skies', officially Stewart Island / Rakiura) is New Zealand's third-largest island, located south of the South Island, across the Foveaux Strait. It is a roughly triangular island with a total land ar ...
. This species is common in lowland and montane forest. Although the largest concentration of trees is now found on the West Coast of the South Island, the biggest trees tend to be in mixed podocarp forest near
Taupō Taupō (), sometimes written Taupo, is a town on the north-eastern shore of Lake Taupō, New Zealand's largest lake, in the central North Island. It is the largest urban area of the Taupō District, and the second-largest urban area in the Wa ...
(e.g., Pureora, Waihaha, and Whirinaki Forests). A typical North Island habitat is in the Hamilton Ecological District, where '' Fuscospora truncata'' and rimu form the
overstory In biology, the canopy is the aboveground portion of a plant cropping or crop, formed by the collection of individual plant crowns. In forest ecology, canopy also refers to the upper layer or habitat zone, formed by mature tree crowns a ...
. Associate ferns on the forest floor are ''
Blechnum discolor ''Lomaria discolor'', synonym ''Blechnum discolor'', commonly called crown fern (Māori: piupiu), is a species of fern in the family Blechnaceae. This species is endemic to New Zealand New Zealand ( mi, Aotearoa ) is an island country i ...
'', '' Blechnum filiforme'', '' Asplenium flaccidum'' and '' Hymenophyllum demissum''. An 800-year-old rimu tree can be seen at the Otari-Wilton's Bush in Wellington.


Description

Rimu is a slow-growing tree, eventually attaining a height of up to 50 m, although most surviving large trees are 20 to 35 m tall. It typically appears as an emergent from mixed broadleaf
temperate rainforest Temperate rainforests are coniferous or broadleaf forests that occur in the temperate zone and receive heavy rain. Temperate rain forests occur in oceanic moist regions around the world: the Pacific temperate rain forests of North American Pac ...
, although there are almost pure stands (especially on the west coast of the
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman S ...
). There are historical accounts of exceptionally tall trees, 61 m, from dense forest near
National Park A national park is a natural park in use for conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state declares or owns. Although individua ...
in the central North Island, now destroyed. Its lifespan is approximately 800 to 900 years. The straight trunk of the rimu is generally 1.5 m in diameter, but may be larger in old or very tall specimens. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
are spirally arranged, awl-shaped, up to 7 mm long on juvenile plants, and 1 mm wide; and 2 to 3 mm long on mature trees. It is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with male and female cones on separate trees; the seeds take 15 months to mature after
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, ...
. The mature cones comprise a swollen red fleshy scale six to ten mm long bearing one (rarely two) apical
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s 4 mm long. The seeds are dispersed by birds which eat the fleshy scale and pass the seed on in their droppings; the cones are an important food resource for some species, particularly the kakapo, whose breeding cycle has been linked to the fruiting cycle of the tree.


Uses

Māori originally used the resinous heartwood of rimu (called māpara or kāpara) for wooden items such as heru (combs) and fernroot beaters. Historically, rimu and other native trees such as
kauri ''Agathis'', commonly known as kauri or dammara, is a genus of 22 species of evergreen tree. The genus is part of the ancient conifer family Araucariaceae, a group once widespread during the Jurassic and Cretaceous periods, but now largely re ...
, matai and
totara ''Podocarpus totara'' (; from the Maori-language ; the spelling "totara" is also common in English) is a species of podocarp tree endemic to New Zealand. It grows throughout the North Island and northeastern South Island in lowland, montane an ...
were the main sources of
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
for New Zealand, including
furniture Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating (tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks). F ...
and
house A house is a single-unit residential building. It may range in complexity from a rudimentary hut to a complex structure of wood, masonry, concrete or other material, outfitted with plumbing, electrical, and heating, ventilation, and air condi ...
construction. However, many of New Zealand's original stands of rimu have been destroyed, and recent government policies forbid the felling of rimu in public forests, though allowing limited logging on private land. ''
Pinus radiata ''Pinus radiata'' ( syn. ''Pinus insignis''), the Monterey pine, insignis pine or radiata pine, is a species of pine native to the Central Coast of California and Mexico (Guadalupe Island and Cedros island). It is an evergreen conifer in the ...
'' has now replaced rimu in most industries, although rimu remains popular for the production of high quality wooden furniture. There is also limited recovery of stump and root wood, from trees felled many years before, for use in making bowls and other wood turned objects. The inner bark can also be used to treat burns and cuts.


In cultivation

Although slow to establish, with a long juvenile period and fairly high moisture requirements, rimu is widely grown as an ornamental tree in New Zealand. It is attractive at all growth stages, usually quite narrow when young, then developing into a broader tree with weeping branches before finally progressing to its more upright adult form. While rimu does exhibit some variation in the wild, garden cultivars are largely unknown, except for one recent introduction, 'Charisma', which is a compact, golden-foliaged form.


References


External links

* {{Taxonbar, from=Q382469 cupressinum Trees of New Zealand Trees of mild maritime climate Least concern plants Plants used in traditional Māori medicine