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The levator ani is a broad, thin
muscle Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscle ...
group, situated on either side of the
pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
. It is formed from three muscle components: the pubococcygeus, the iliococcygeus, and the puborectalis. It is attached to the inner surface of each side of the
lesser pelvis The pelvic cavity is a body cavity that is bounded by the bones of the pelvis. Its oblique roof is the pelvic inlet (the superior opening of the pelvis). Its lower boundary is the pelvic floor. The pelvic cavity primarily contains the reproduct ...
, and these unite to form the greater part of the
pelvic floor The pelvic floor or pelvic diaphragm is composed of muscle fibers of the levator ani, the coccygeus muscle, and associated connective tissue which span the area underneath the pelvis. The pelvic diaphragm is a muscular partition formed by the lev ...
. The
coccygeus muscle The coccygeus muscle or ischiococcygeus is a muscle of the pelvic floor, located posterior to levator ani and anterior to the sacrospinous ligament. Structure The coccygeus muscle is posterior to levator ani and anterior to the sacrospinous lig ...
completes the pelvic floor, which is also called the ''pelvic diaphragm''. It supports the
viscera In biology, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to act together in a ...
in the
pelvic cavity The pelvic cavity is a body cavity that is bounded by the bones of the pelvis. Its oblique roof is the pelvic inlet (the superior opening of the pelvis). Its lower boundary is the pelvic floor. The pelvic cavity primarily contains the reproduct ...
, and surrounds the various structures that pass through it. The levator ani is the main
pelvic floor The pelvic floor or pelvic diaphragm is composed of muscle fibers of the levator ani, the coccygeus muscle, and associated connective tissue which span the area underneath the pelvis. The pelvic diaphragm is a muscular partition formed by the lev ...
muscle and painfully contracts during
vaginismus Vaginismus is a condition in which involuntary muscle spasm interferes with vaginal intercourse or other penetration of the vagina. This often results in pain with attempts at sex. Often, it begins when vaginal intercourse is first attempted. ...
. It also contracts rhythmically during
orgasm Orgasm (from Greek , ; "excitement, swelling") or sexual climax is the sudden discharge of accumulated sexual excitement during the sexual response cycle, resulting in rhythmic, involuntary muscular contractions in the pelvic region charac ...
.


Structure

The levator ani is made up of 3 parts: * Iliococcygeus muscle * Pubococcygeus muscle * Puborectalis muscle The iliococcygeus arises from the inner side of the
ischium The ischium () form ...
(the lower and back part of the
hip bone The hip bone (os coxae, innominate bone, pelvic bone or coxal bone) is a large flat bone, constricted in the center and expanded above and below. In some vertebrates (including humans before puberty) it is composed of three parts: the ilium, ischi ...
) and from the posterior part of the tendinous arch of the
obturator fascia The obturator fascia, or fascia of the internal obturator muscle, covers the pelvic surface of that muscle and is attached around the margin of its origin. Above, it is loosely connected to the back part of the arcuate line, and here it is conti ...
, and is attached to the
coccyx The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...
and
anococcygeal body The anococcygeal body (anococcygeal ligament, or anococcygeal raphe) is a fibrous median raphe in the floor of the pelvis, which extends between the coccyx and the margin of the anus. It is composed of fibers of the levator ani muscle that unite ...
; it is usually thin, and may be absent, or be largely replaced by fibrous tissue. An accessory
slip Slip or SLIP may refer to: Science and technology Biology * Slip (fish), also known as Black Sole * Slip (horticulture), a small cutting of a plant as a specimen or for grafting * Muscle slip, a branching of a muscle, in anatomy Computing and ...
at its posterior part is sometimes named the ''iliosacralis''. The pubococcygeus muscle has medial fibres forming the pubovaginalis in the female, and the puboprostaticus in the male.


Origin and insertion

The levator ani arises, in front, from the posterior surface of the
superior pubic ramus In vertebrates, the pubic region ( la, pubis) is the most forward-facing (ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior ra ...
lateral to the
symphysis A symphysis (, pl. symphyses) is a fibrocartilaginous fusion between two bones. It is a type of cartilaginous joint, specifically a secondary cartilaginous joint. # A symphysis is an amphiarthrosis, a slightly movable joint. # A growing together ...
; behind, from the inner surface of the spine of the
ischium The ischium () form ...
; and between these two points, from the
obturator fascia The obturator fascia, or fascia of the internal obturator muscle, covers the pelvic surface of that muscle and is attached around the margin of its origin. Above, it is loosely connected to the back part of the arcuate line, and here it is conti ...
. Posteriorly, this fascial origin corresponds, more or less closely, with the tendinous arch of the pelvic fascia, but in front, the muscle arises from the fascia at a varying distance above the arch, in some cases reaching nearly as high as the canal for the obturator vessels and nerve. The fibers pass downward and backward to the middle line of the floor of the pelvis; the most posterior are inserted into the side of the last two segments of the
coccyx The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...
; those placed more anteriorly unite with the muscle of the opposite side, in a median fibrous ridge called the
anococcygeal body The anococcygeal body (anococcygeal ligament, or anococcygeal raphe) is a fibrous median raphe in the floor of the pelvis, which extends between the coccyx and the margin of the anus. It is composed of fibers of the levator ani muscle that unite ...
or raphe, which extends between the coccyx and the margin of the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, de ...
. The middle fibers are inserted into the side of the
rectum The rectum is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. The adult human rectum is about long, and begins at the rectosigmoid junction (the end of the sigmoid colon) at the le ...
, blending with the fibers of the
sphincter A sphincter is a circular muscle that normally maintains constriction of a natural body passage or orifice and which relaxes as required by normal physiological functioning. Sphincters are found in many animals. There are over 60 types in the hum ...
muscles; lastly in the male, the anterior fibers descend upon the side of the prostate to unite beneath it with the muscle of the opposite side, joining with the fibers of the
external anal sphincter The external anal sphincter (or sphincter ani externus ) is a flat plane of skeletal muscle fibers, elliptical in shape and intimately adherent to the skin surrounding the margin of the anus. Anatomy The external anal sphincter measures about 8 t ...
and
transverse perineal muscles The transverse perineal muscles (transversus perinei) are the superficial and the deep transverse perineal muscles. Superficial transverse perineal ...
, at the central tendinous point of the
perineum The perineum in humans is the space between the anus and scrotum in the male, or between the anus and the vulva in the female. The perineum is the region of the body between the pubic symphysis (pubic arch) and the coccyx (tail bone), includin ...
. The anterior portion is occasionally separated from the rest of the muscle by connective tissue. From this circumstance, as well as from its peculiar relation with the
prostate The prostate is both an accessory gland of the male reproductive system and a muscle-driven mechanical switch between urination and ejaculation. It is found only in some mammals. It differs between species anatomically, chemically, and physio ...
, which it supports as in a sling, it has been described as a distinct muscle, under the name of ''levator prostatæ''. In the female the anterior fibers of the levator ani descend upon the side of the
vagina In mammals, the vagina is the elastic, muscular part of the female genital tract. In humans, it extends from the vestibule to the cervix. The outer vaginal opening is normally partly covered by a thin layer of mucosal tissue called the hymen ...
.


Innervation

The levator ani muscles are mostly innervated by the
pudendal nerve The pudendal nerve is the main nerve of the perineum. It carries sensation from the external genitalia of both sexes and the skin around the anus and perineum, as well as the motor supply to various pelvic muscles, including the male or fema ...
,
perineal nerve The perineal nerve is a nerve of the pelvis. It arises from the pudendal nerve in the pudendal canal. It gives superficial branches to the skin, and a deep branch to muscles. It supplies the skin and muscles of the perineum. Its latency is tes ...
and acting together.


Variation

In addition,
sacral spinal nerves A spinal nerve is a mixed nerve, which carries motor, sensory, and autonomic signals between the spinal cord and the body. In the human body there are 31 pairs of spinal nerves, one on each side of the vertebral column. These are grouped into the ...
( S3, S4) innervate the muscles directly as well (in ~70% of people). Sometimes (in ~40% of people) the inferior rectal nerve innervates the levator ani muscles independently of the pudendal nerve.


Pubococcygeus muscle

The pubococcygeus muscle or PC muscle is a hammock-like muscle, found in both sexes, that stretches from the
pubic bone In vertebrates, the pubic region ( la, pubis) is the most forward-facing (ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior r ...
to the
coccyx The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...
(tail bone) forming the
floor A floor is the bottom surface of a room or vehicle. Floors vary from simple dirt in a cave to many layered surfaces made with modern technology. Floors may be stone, wood, bamboo, metal or any other material that can support the expected load ...
of the
pelvic cavity The pelvic cavity is a body cavity that is bounded by the bones of the pelvis. Its oblique roof is the pelvic inlet (the superior opening of the pelvis). Its lower boundary is the pelvic floor. The pelvic cavity primarily contains the reproduct ...
and supporting the pelvic organs.


Structure

The pubococcygeus arises from the back of the pubis and from the anterior part of the
obturator fascia The obturator fascia, or fascia of the internal obturator muscle, covers the pelvic surface of that muscle and is attached around the margin of its origin. Above, it is loosely connected to the back part of the arcuate line, and here it is conti ...
, and is directed backward almost horizontally along the side of the
anal canal The anal canal is the part that connects the rectum to the anus, located below the level of the pelvic diaphragm. It is located within the anal triangle of the perineum, between the right and left ischioanal fossa. As the final functional segme ...
toward the
coccyx The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other ...
and
sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
, to which it finds attachment. Between the termination of the vertebral column and the anus, the two pubococcygeus muscles come together and form a thick, fibromuscular layer lying on the raphe (ridge) or (
anococcygeal body The anococcygeal body (anococcygeal ligament, or anococcygeal raphe) is a fibrous median raphe in the floor of the pelvis, which extends between the coccyx and the margin of the anus. It is composed of fibers of the levator ani muscle that unite ...
) formed by the iliococcygei. The greater part of this muscle is inserted into the coccyx and into the last one or two pieces of the sacrum.


Variation

This insertion into the vertebral column is, however, not accepted by all observers.


Function

The pubococcygeus muscle controls urine flow and contracts during
orgasm Orgasm (from Greek , ; "excitement, swelling") or sexual climax is the sudden discharge of accumulated sexual excitement during the sexual response cycle, resulting in rhythmic, involuntary muscular contractions in the pelvic region charac ...
as well as assisting in male
ejaculation Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential component ...
. It also aids in childbirth as well as core stability. A strong pubococcygeus muscle has also been linked to a reduction in
urinary incontinence Urinary incontinence (UI), also known as involuntary urination, is any uncontrolled leakage of urine. It is a common and distressing problem, which may have a large impact on quality of life. It has been identified as an important issue in geria ...
and proper positioning of the baby's head during childbirth.


Kegel exercises

The
Kegel exercise Kegel exercise, also known as pelvic-floor exercise, involves repeatedly contracting and relaxing the muscles that form part of the pelvic floor, now sometimes colloquially referred to as the "Kegel muscles". The exercise can be performed many ti ...
s are a series of voluntary contractions of all the perineal muscles. Such movement is done in an effort to strengthen all the striated muscles in the perineum's area. They are often referred to simply as "kegels", named after their founder, Dr.
Arnold Kegel Arnold Henry Kegel (February 21, 1894 – March 1, 1972) was an American gynecologist who invented the Kegel perineometer (an instrument for measuring the strength of voluntary contractions of the pelvic floor muscles) and Kegel exercises (s ...
. These exercises also serve to contract, among others, the
ischiocavernosus The ischiocavernosus muscle (erectores penis ''or'' erector clitoridis in older texts) is a muscle just below the surface of the perineum, present in both men and women. Structure It arises by tendinous and fleshy fibers from the inner surface of ...
,
bulbospongiosus The bulbospongiosus muscle (bulbocavernosus in older texts) is one of the superficial muscles of the perineum. It has a slightly different origin, insertion and function in males and females. In males, it covers the bulb of the penis. In fema ...
, and
cremaster muscle The cremaster muscle is a paired structure made of thin layers of striated and smooth muscle that covers the testis and the spermatic cord in human males. It consists of the lateral and medial parts. Cremaster is an involuntary muscle, responsi ...
in men, as voluntary contraction of the pubococcygeus muscle also engages the
cremasteric reflex The cremasteric reflex is a superficial (i.e., close to the skin's surface) reflex observed in human males. This reflex is elicited by lightly stroking or poking the superior and medial (inner) part of the thigh—regardless of the direction of ...
, which lifts the testicles up, although this does not occur in all men. Kegel exercises have been prescribed to ameliorate
erectile dysfunction Erectile dysfunction (ED), also called impotence, is the type of sexual dysfunction in which the penis fails to become or stay erect during sexual activity. It is the most common sexual problem in men.Cunningham GR, Rosen RC. Overview of male ...
due to venous leakage and to help men control
premature ejaculation Premature ejaculation (PE) occurs when a man expels semen (and most likely experiences orgasm) soon after beginning sexual activity, and with minimal penile stimulation. It has also been called ''early ejaculation'', ''rapid ejaculation'', ''ra ...
and to treat urinary incontinence in both sexes.


Puborectalis muscle

The fibers that form a
sling sling may refer to: Places * Sling, Anglesey, Wales * Sling, Gloucestershire, England, a small village in the Forest of Dean People with the name * Otto Šling (1912–1952), repressed Czech communist functionary Arts, entertainment, and media ...
looping around the
rectum The rectum is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. The adult human rectum is about long, and begins at the rectosigmoid junction (the end of the sigmoid colon) at the le ...
are named the puborectalis. They arise from the lower part of the
pubic symphysis The pubic symphysis is a secondary cartilaginous joint between the left and right superior rami of the pubis of the hip bones. It is in front of and below the urinary bladder. In males, the suspensory ligament of the penis attaches to the pubic ...
, and from the
superior fascia of the urogenital diaphragm Superior may refer to: *Superior (hierarchy), something which is higher in a hierarchical structure of any kind Places *Superior (proposed U.S. state), an unsuccessful proposal for the Upper Peninsula of Michigan to form a separate state *Lake ...
. The origin of the puborectalis fibers is at the posterior surface of the pubis while their insertion is at the midline sling posterior to the rectum. The muscle band is innervated by perineal branches of the S3 and S4 nerve roots. They meet with the corresponding fibers of the opposite side around the lower part of the rectum, and form for it a strong sling. Relaxation increases the angle between rectum and anus, allowing defecation in conjunction with relaxation of the
internal Internal may refer to: *Internality as a concept in behavioural economics *Neijia, internal styles of Chinese martial arts *Neigong or "internal skills", a type of exercise in meditation associated with Daoism *''Internal (album)'' by Safia, 2016 ...
and
external anal sphincter The external anal sphincter (or sphincter ani externus ) is a flat plane of skeletal muscle fibers, elliptical in shape and intimately adherent to the skin surrounding the margin of the anus. Anatomy The external anal sphincter measures about 8 t ...
s. Levator ani relaxation and rectal emptying is facilitated by anorectal straightening during squatting.


Function

The levator ani muscles are responsible for "wagging" the tail in
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, ...
ed
quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor ...
s. These muscles are not as strong in the human, as tail-wagging is more demanding than the support function that the muscles serve in humans.


Clinical significance


Levator ani syndrome

Levator ani syndrome Levator ani syndrome is a condition characterized by burning pain or tenesmus of the rectal or perineal area, caused by spasm of the levator ani muscle.Levator Syndrome, by Parswa Ansari, MD 7/2014, Merck Manuals The genesis of the syndrome is unk ...
is episodic
rectal The rectum is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. The adult human rectum is about long, and begins at the rectosigmoid junction (the end of the sigmoid colon) at the le ...
pain caused by spasm of the levator ani muscle.Levator Syndrome, by Parswa Ansari, MD 7/2014, Merck Manuals


Levator ani avulsion

https://pubmed.ncbi.nlm.nih.gov/34245656/


Additional images

File:Gray236.png, Right hip bone. Internal surface. File:Gray402.png, Coronal section of pelvis, showing arrangement of fasciæ. Viewed from behind. File:1116 Muscle of the Male Perineum.png, Muscles of male perineum. File:Levator ani.png File:Gray539.png, The arteries of the pelvis. File:Gray837.png, Sacral plexus of the right side. File:Gray1076.png, Iliac colon, sigmoid or pelvic colon, and rectum seen from the front, after removal of pubic bones and bladder. File:Gray1077.png, The posterior aspect of the rectum exposed by removing the lower part of the sacrum and the coccyx. File:Gray1136.png, Male pelvic organs seen from right side. File:Human anus-en.svg, Anatomy of the human anus.


See also

*
Bulbospongiosus muscle The bulbospongiosus muscle (bulbocavernosus in older texts) is one of the superficial muscles of the perineum. It has a slightly different origin, insertion and function in males and females. In males, it covers the bulb of the penis. In femal ...
*
Coccygectomy Coccygectomy is a surgical procedure in which the coccyx or tailbone is removed. It is considered a required treatment for sacrococcygeal teratoma and other germ cell tumors arising from the coccyx. Coccygectomy is the treatment of last resort for ...


References


External links

* —"Muscles of the female superficial perineal pouch." * —"Muscles of the male superficial perineal pouch." * —"Muscles of the Pelvic Diaphragm" * * * * * () * (, )
Merck Manual article on levator ani syndrome
{{DEFAULTSORT:Levator Ani Muscles of the torso Pelvis Sexual anatomy