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The philtrum ( la, philtrum from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
''phíltron,'' lit. "love charm"), or medial cleft, is a vertical indentation in the middle area of the upper lip, common to therian mammals, extending in humans from the
nasal septum The nasal septum () separates the left and right airways of the nasal cavity, dividing the two nostrils. It is depressed by the depressor septi nasi muscle. Structure The fleshy external end of the nasal septum is called the columella or co ...
to the
tubercle of the upper lip The lips are the visible body part at the mouth of many animals, including humans. Lips are soft, movable, and serve as the opening for food intake and in the articulation of sound and speech. Human lips are a tactile sensory organ, and can be ...
. Together with a glandular
rhinarium The rhinarium (New Latin, "belonging to the nose"; plural: rhinaria) is the furless skin surface surrounding the external openings of the nostrils in many mammals. Commonly it is referred to as the tip of the ''snout'', and breeders of cats and ...
and slit-like
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
s, it is believed to constitute the primitive condition for at least therian mammals. Monotremes lack a philtrum, though this could be due to the specialised, beak-like jaws in living species.


Function

In most mammals, the philtrum is a narrow groove that may carry dissolved odorants from the
rhinarium The rhinarium (New Latin, "belonging to the nose"; plural: rhinaria) is the furless skin surface surrounding the external openings of the nostrils in many mammals. Commonly it is referred to as the tip of the ''snout'', and breeders of cats and ...
or nose pad to the vomeronasal organ via ducts inside the mouth. For humans and most primates, the philtrum survives only as a
vestigial Vestigiality is the retention, during the process of evolution, of genetically determined structures or attributes that have lost some or all of the ancestral function in a given species. Assessment of the vestigiality must generally rely on co ...
medial depression between the nose and upper lip. The human philtrum, bordered by ridges, is also known as the ''infranasal depression'', but has no apparent function. That may be because most higher primates rely more on vision than on smell.
Strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and southeast Asia. Colle ...
primates, such as
lemur Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are wet-nosed primates of the superfamily Lemuroidea (), divided into 8 families and consisting of 15 genera and around 100 existing species. They are endemic to the island of Madagas ...
s, still retain the philtrum and the rhinarium, unlike monkeys and apes.


Development

In humans, the philtrum is formed where the nasomedial and maxillary processes meet during embryonic development. When these processes fail to fuse fully, a
cleft lip A cleft lip contains an opening in the upper lip that may extend into the nose. The opening may be on one side, both sides, or in the middle. A cleft palate occurs when the palate (the roof of the mouth) contains an opening into the nose. The te ...
may result.


Variation

A flattened or smooth philtrum may be a symptom of fetal alcohol syndrome or Prader–Willi syndrome. A study of boys diagnosed with
autism spectrum disorders The autism spectrum, often referred to as just autism or in the context of a professional diagnosis autism spectrum disorder (ASD) or autism spectrum condition (ASC), is a neurodevelopmental condition (or conditions) characterized by difficulti ...
found that a broader than average philtrum is one of a cluster of physical attributes associated with autism.


Society and culture

In
Jewish tradition Jews ( he, יְהוּדִים, , ) or Jewish people are an ethnoreligious group and nation originating from the Israelites Israelite origins and kingdom: "The first act in the long drama of Jewish history is the age of the Israelites"" ...
, each embryo has an angel teaching it all of the wisdom in the world while it is in utero. The angel lightly taps the infant's upper lip before birth to prevent the infant from revealing the secrets of the universe; the infant then forgets the Torah it has been taught. Some believers of the story speculate that this is the cause of the philtrum, but it does not have a basis in traditional Jewish texts. In ''
Key Largo Key Largo ( es, Cayo Largo) is an island in the upper Florida Keys archipelago and is the largest section of the keys, at long. It is one of the northernmost of the Florida Keys in Monroe County, and the northernmost of the keys connected by ...
'' (1948), Frank McCloud ( Humphrey Bogart) tells a fairy tale, saying that, before birth, the soul knows all the secrets of heaven, but at birth an angel presses a fingertip just above one's lip, which seals us to silence. This is also cited in the Stephen King short story " Afterlife". In the movie '' Mr. Nobody'', unborn infants are said to have knowledge of all past and future events. As an unborn infant is about to be sent to its mother, the "Angels of Oblivion" lightly tap its upper lip, whereupon the unborn infant forgets everything it knows. The movie follows the life story of an infant whose lip had not been tapped. In the book ''Prince Ombra'' by Roderick MacLeish, the "cleft on our upper lips" is attributed to being hushed by a "cavern angel" just before we are born. In Philippine mythology the enchanted creature ''
diwata ''Anito'', also spelled ''anitu'', refers to ancestor spirits, nature spirits, and deities in the indigenous Philippine folk religions from the precolonial age to the present, although the term itself may have other meanings and associa ...
'' (or encantado) has smooth skin, with no wrinkles even at the joints, and no philtrum. In '' Les Misérables'' by Victor Hugo, translated by Isabel F. Hapgood, Fantine's philtrum is described thus, "in the very characteristic interval which separates the base of the nose from the upper lip, she had that imperceptible and charming fold, a mysterious sign of chastity, which makes Barberousse fall in love with a Diana found in the treasures of Iconia." Book Third—In The Year 1817, Chapter III—Four And Four.


See also

*
Cupid's bow The Cupid's bow is a facial feature where the double curve of a human upper lip is said to resemble the bow of Cupid, the Roman god of erotic love. The peaks of the bow coincide with the philtral columns giving a prominent bow appearance to the ...
* Intermaxillary segment *
Toothbrush moustache The toothbrush moustache is a style of moustache in which the sides are vertical (or nearly vertical) rather than tapered, giving the hairs the appearance of the bristles on a toothbrush that are attached to the nose. It was made famous by such ...
(philtrum moustache) *
Philtrum piercing A philtrum piercing, nicknamed a medusa piercing, is an upper lip piercing placed in the philtrum, directly under the septum of the nose. It is typically pierced using a labret A labret is a form of body piercing. Taken literally, it is any typ ...


References

{{Authority control Mammal anatomy Facial features Lips Primate anatomy Horse anatomy Rodent anatomy Dog anatomy