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The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a Palearctic hare that is largely adapted to polar and mountainous habitats.


Evolution

The mountain hare arose during the Late Pleistocene; there is evidence that its range expanded during glaciations into southern Europe, with populations of
Iberian hare The Granada hare (''Lepus granatensis''), also known as the Iberian hare, is a hare species that can be found on the Iberian Peninsula and on the island of Majorca. Subspecies Three subspecies of the Granada hare are known, which vary in colour ...
(''Lepus granatensis''),
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
(''L. europaeus)'' and broom hare (''L. castroviejoi'') in northern Iberia harboring mitochondrial haplotypes from the mountain hare. During the Late Pleistocene to
Early Holocene In the geologic time scale, the Greenlandian is the earliest age or lowest stage of the Holocene Epoch or Series, part of the Quaternary. Beginning in 11,650 BP (9701 BCE or 300 HE) and ending 8,276 BP (6237 BCE or 3764 HE), it is the earlie ...
, populations of mountain hare in Russia grew at least 10% larger than any living population today. This population has been classified as a distinct species ''Lepus tanaiticus'', but is now generally considered a prehistoric morphotype of the living mountain hare.


Distribution

This species is distributed from Fennoscandia to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive region, geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a ...
; in addition, isolated mountain populations occur in the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Swi ...
,
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
, the
Baltics The Baltic states, et, Balti riigid or the Baltic countries is a geopolitical term, which currently is used to group three countries: Estonia, Latvia, and Lithuania. All three countries are members of NATO, the European Union, the Eurozone, ...
, northeastern
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
, and Hokkaidō. In
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
, the Irish hare (''L. t. hibernicus'') lives on lowland pastures, coastal grasslands, and salt marshes, not just in the mountains. The mountain hare has also been introduced to
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...
, Shetland, Orkney, the
Isle of Man ) , anthem = "O Land of Our Birth" , image = Isle of Man by Sentinel-2.jpg , image_map = Europe-Isle_of_Man.svg , mapsize = , map_alt = Location of the Isle of Man in Europe , map_caption = Location of the Isle of Man (green) in Europe ...
, the
Peak District The Peak District is an upland area in England at the southern end of the Pennines. Mostly in Derbyshire, it extends into Cheshire, Greater Manchester, Staffordshire, West Yorkshire and South Yorkshire. It includes the Dark Peak, where moorla ...
, Svalbard, Kerguelen Islands, Crozet Islands, and the
Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...
. In the Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on geographic region and season.


Description

The mountain hare is a large species, though it is slightly smaller than the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
. It grows to a length of , with a tail of , and a mass of , females being slightly heavier than males. They can live for up to 12 years. In summer, for all populations of mountain hares, the coat is various shades of brown. In preparation for winter most populations moult into a white (or largely white)
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
. The tail remains completely white all year round, distinguishing the mountain hare from the
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
(''Lepus europaeus''), which has a black upper side to the tail. The subspecies ''Lepus timidus hibernicus'' (the Irish mountain hare) is smaller in size and stays brown all year. The Irish hare may also have a "golden" variation, particularly those found on Rathlin Island. In the Faroe Islands, mountain hares turn grey in the winter instead of white. The winter-grey colour may be caused by downregulation of the agouti hair cycle isoform in the autumn moult.


Behaviour

The diet of the mountain hare varies from region to region. It seems to be somewhat dependent on the particular habitat in which the population under study lives. For example, in northern Scandinavia, where snow may blanket the ground for many months, the hares may feed on twigs and bark. In areas where snowfall is rare, such as Ireland,
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
may form the bulk of the diet. Given a choice, mountain hares in Scotland and Ireland seem to prefer grazing (feeding on grasses). For mountain hares on a coastal
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
environment in Ireland, grasses constituted over 90% of their diet. This was higher than the percentage of grass in the diet of the
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has b ...
(''Oryctolagus cuniculus'') that inhabited the same environment. The mountain hare is regionally the favourite prey of the
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds of ...
and may additionally be preyed on by
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and femal ...
s and red foxes.
Stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
s may prey on young hares. In northern parts of Finland, Norway, and Sweden, the mountain hare and the European hare compete for habitat. The European hare, being larger, is usually able to drive away the mountain hare, but is less adapted for living in snowy regions; its feet are smaller and its winter fur is a mixture of white and brown. While this winter fur is actually a very good camouflage in the coastal regions of Finland where the snow covers the shrubs only a short time, the mountain hare is better adapted for the snowier conditions of the inland areas. The two may occasionally cross. The
Arctic hare The Arctic hare (''Lepus arcticus'') is a species of hare highly adapted to living in the Arctic tundra and other icy biomes. The Arctic hare survives with shortened ears and limbs, a small nose, fat that makes up close to 20% of its body, and a ...
(''Lepus arcticus'') was once considered a subspecies of the mountain hare, but it is now regarded as a separate species. Similarly, some scientists believe that the Irish hare should be regarded as a separate species. Fifteen subspecies are currently recognised.


Human impact

In the European Alps, the mountain hare lives at elevations from 700 to 3800 m, depending on biographic region and season. The development of alpine winter tourism has increased rapidly since the last few decades of the 20th century, resulting in expansion of ski resorts, growing visitor numbers, and a huge increase in all forms of snow sport activities. A 2013 study looking at stress events and the response of mountain hares to disturbance concluded that those hares living in areas of high winter recreational activities showed changes in physiology and behaviour that demanded additional energy input at a time when access to food resources is restricted by snow. It recommended ensuring that forests inhabited by mountain hares were kept free of tourist development, and that new skiing areas should be avoided in mountain hare habitat, and that existing sites should not be expanded. In August 2016, the Scottish animal welfare charity
OneKind OneKind is a Advocacy group, campaigning animal welfare charity based in Edinburgh and operating in Scotland, UK and as part of the Eurogroup for Animals. OneKind exists to end cruelty to all animals and improve their lives. Background The org ...
launched a campaign on behalf of the mountain hare, as a way of raising awareness of mountain hare culls taking place across the country and in garnering public support for the issue. Mountain hares are routinely shot in the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland S ...
both as part of paid hunting "tours" and by gamekeepers managing red grouse populations (who believe that mountain hares can be vectors of diseases that affect the birds). Much of this activity is secretive, but investigations have revealed that tens of thousands of hares are being culled every year. The campaign, which urges people to proclaim, "We Care For The Mountain Hare", will culminate with the charity urging the Scottish government to legislate against commercial hunting and culling of the iconic Scottish species. The campaign has revealed widespread public support for a ban on hare hunting in Scotland. On May 17, 2020, MSPs voted to ban the unlicensed culling of mountain hares and grant them protected species status within Scotland after a petition started by Green MSP
Alison Johnstone Alison Johnstone (born 11 October 1965) is a Scottish politician who has served as the Presiding Officer of the Scottish Parliament since 2021. Elected as a member of the Scottish Greens, she relinquished her party affiliation on becoming Presi ...
gathered over 22,000 signatures. In 2021, the People's Trust for Endangered Species funded a survey of mountain hare populations in the UK's Peak District after concerns about the viability of the isolated population, believed to be as low as 2,500. The trust believes climate change is a threat to long-term survival of the Peak District population, which was introduced to the area in the 1870s.


References


External links


Irish Hare Initiative
{{Taxonbar, from=Q180035 Lepus Mammals of Asia Mammals of Europe Mammals of China Mammals of Japan Mammals of Central Asia Mammals of Russia Arctic land animals Mammals described in 1758 Taxa named by Carl Linnaeus National symbols of the Republic of Ireland Mammals of Mongolia