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''Dimocarpus longan'', commonly known as the longan () and dragon's eye, is a tropical tree species that produces edible fruit. It is one of the better-known tropical members of the soapberry family
Sapindaceae The Sapindaceae are a family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1858 accepted species. Examples include horse chestnut, maples, ackee and lychee. The Sapindaceae occur in tempe ...
, to which the
lychee Lychee (US: ; UK: ; ''Litchi chinensis''; ) is a monotypic taxon and the sole member in the genus ''Litchi'' in the soapberry family, ''Sapindaceae''. It is a tropical tree native to Southeast and Southwest China (the Guangdong, Fujian, Y ...
and rambutan also belong. The fruit of the longan is similar to that of the lychee, but less aromatic in taste. It is native to tropical Asia and China. The longan (from
Cantonese Cantonese ( zh, t=廣東話, s=广东话, first=t, cy=Gwóngdūng wá) is a language within the Chinese (Sinitic) branch of the Sino-Tibetan languages originating from the city of Guangzhou (historically known as Canton) and its surrounding are ...
''lùhng-ngáahn'' , literally 'dragon eye'), is so named because it resembles an eyeball when its fruit is shelled (the black seed shows through the translucent flesh like a
pupil The pupil is a black hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black ...
and iris). The seed is small, round and hard, and of an enamel-like, lacquered black. The fully ripened, freshly harvested fruit has a bark-like shell, thin, and firm, making the fruit easy to peel by squeezing the pulp out as if one were "cracking" a sunflower seed. When the shell has more moisture content and is more tender, the fruit becomes less convenient to shell. The tenderness of the shell varies due to either premature harvest, variety, weather conditions or storage conditions.


Subspecies

Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants b ...
lists: * ''D. longan'' var. ''echinatus'' Leenhouts (Borneo, Philippines) * ''D. longan'' var. ''longetiolatus'' Leenhouts (Viet Nam) * ''D. longan'' subsp. ''malesianus'' Leenh. (widespread SE Asia) * ''D. longan'' var. ''obtusus'' (Pierre) Leenh. (Indo-China)


Tree description

Depending upon climate and soil type the tree may grow to over in height, but it typically stands in height and the
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
is round. The trunk is thick with corky bark. The branches are long and thick, typically drooping. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ...
are oblong and blunt-tipped, usually long and wide. The leaves are
pinnately Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
compounded and alternate. There are 6 to 9 pairs of leaflets per leaf and the upper surface is wavy and a dark, glossy-green. The longan tree produces light-yellow
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s at the end of branches. The inflorescence is commonly called a
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are ...
and are long, and widely branched. The small
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...
s have 5 to 6
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined ...
s and
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
s that are brownish-yellow. The flower has a two-lobed
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pis ...
and 8
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
. There are three flower types, distributed throughout the panicle; staminate (functionally male), pistillate (functionally female), and
hermaphroditic In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many taxonomic groups of animals (mostly invertebrates) do not have sepa ...
flowers. Flowering occurs as a progression. The fruit hangs in drooping clusters that are circular and about wide. The peel is tan, thin, and leathery with tiny hairs. The flesh is translucent, and the
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
is large and black with a circular white spot at the base. This gives the illusion of an eye. The flesh has a musky, sweet taste, which can be compared to the flavor of lychee fruit. The longan tree is somewhat sensitive to
frost Frost is a thin layer of ice on a solid surface, which forms from water vapor in an above- freezing atmosphere coming in contact with a solid surface whose temperature is below freezing, and resulting in a phase change from water vapor (a g ...
. Longan trees prefer sandy soil. While the species prefers temperatures that do not typically fall below , it can withstand brief temperature drops to about . Longan trees prefer sandy soil with mild levels of acidity and
organic matter Organic matter, organic material, or natural organic matter refers to the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have ...
. Longans usually bear fruit slightly later than lychees. The wild longan population have been decimated considerably by large-scale logging in the past, and the species used to be listed as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolog ...
. If left alone, longan tree stumps will resprout and the listing was upgraded to
Near Threatened A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify f ...
in 1998. Recent field data are inadequate for a contemporary IUCN assessment.


History

The longan is believed to originate from the mountain range between
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
and southern China. Other reported origins include India,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
,
upper Myanmar Upper Myanmar ( my, အထက်မြန်မာပြည်, also called Upper Burma) is a geographic region of Myanmar, traditionally encompassing Mandalay and its periphery (modern Mandalay, Sagaing, Magway Regions), or more broadly sp ...
, north Thailand,
Kampuchea Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand ...
(more commonly known as Cambodia), north
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making it ...
and
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the List of islands by area, world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Mainlan ...
. Its earliest record of existence draws back to the
Han dynasty The Han dynasty (, ; ) was an imperial dynasty of China (202 BC – 9 AD, 25–220 AD), established by Liu Bang (Emperor Gao) and ruled by the House of Liu. The dynasty was preceded by the short-lived Qin dynasty (221–207 BC) and a wa ...
in 200 BC. The emperor had demanded lychee and longan trees to be planted in his palace gardens in
Shaanxi Shaanxi (alternatively Shensi, see § Name) is a landlocked province of China. Officially part of Northwest China, it borders the province-level divisions of Shanxi (NE, E), Henan (E), Hubei (SE), Chongqing (S), Sichuan (SW), Gansu (W), Ningx ...
, but the plants failed. Four hundred years later, longan trees flourished in other parts of China like
Fujian Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its ca ...
and
Guangdong Guangdong (, ), alternatively romanized as Canton or Kwangtung, is a coastal province in South China on the north shore of the South China Sea. The capital of the province is Guangzhou. With a population of 126.01 million (as of 2020) ...
, where longan production soon became an industry. Later on, due to immigration and the growing demand for nostalgic foods, the longan tree was officially introduced to Australia in the mid-1800s, Thailand in the late-1800s, and Hawaii and Florida in the 1900s. The warm, sandy-soiled conditions allowed for the easy growth of longan trees. This jump-started the longan industry in these locations. Despite its long success in China, the longan is considered to be a relatively new fruit to the world. It has only been acknowledged outside of China in the last 250 years. The first European acknowledgment of the fruit was recorded by Joao de Loureiro, a Portuguese Jesuit botanist, in 1790. The first entry resides in his collection of works, ''Flora Cochinchinensis''. Currently, longan crops are grown in southern China, Taiwan, northern Thailand, Malaysia, Indonesia, Cambodia, Laos, Vietnam, India, Sri Lanka,
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
, Australia, the United States, and
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It inc ...
. It is also grown in
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
.


Culinary uses

The fruit is sweet, juicy, and succulent in superior agricultural varieties. The seed and the peel are not consumed. Apart from being eaten raw like other fruits, longan fruit is also often used in Asian soups, snacks, desserts, and
sweet-and-sour Sweet and sour is a generic term that encompasses many styles of sauce, cuisine and cooking methods. It is commonly used in East Asia and Southeast Asia, and has been used in England since the Middle Ages. Dickson Wright, Clarissa (2011) ''A Histor ...
foods, either fresh or dried, and sometimes preserved and canned in syrup. The taste is different from lychees; while longan has a drier sweetness similar to dates, lychees are often messily juicy with a more tropical, grape-like sour sweetness. Dried longan are often used in
Chinese cuisine Chinese cuisine encompasses the numerous cuisines originating from China, as well as overseas cuisines created by the Chinese diaspora. Because of the Chinese diaspora and historical power of the country, Chinese cuisine has influenced many o ...
and Chinese sweet dessert soups. In
Chinese food therapy Chinese food therapy (, also called nutrition therapy and dietary therapy) is a mode of dieting rooted in Chinese beliefs concerning the effects of food on the human organism, and centered on concepts such as eating in moderation. Its basic pr ...
and
herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remedi ...
, it is believed to have an effect on relaxation. In contrast with the fresh fruit, which is juicy and white, the flesh of dried longans is dark brown to almost black. Once fermented, it can be made into longan wine.


Nutrition

Raw longan fruit is 83% water, 15%
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s, 1% protein, and contains negligible fat. In a 100 gram reference amount, raw longan supplies 60
calorie The calorie is a unit of energy. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of o ...
s of food energy, 101% of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of healt ...
(DV) of
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) ...
, 12% DV of
riboflavin Riboflavin, also known as vitamin B2, is a vitamin found in food and sold as a dietary supplement. It is essential to the formation of two major coenzymes, flavin mononucleotide and flavin adenine dinucleotide. These coenzymes are involved in ...
, and no other
micronutrient Micronutrients are essential dietary elements required by organisms in varying quantities throughout life to orchestrate a range of physiological functions to maintain health. Micronutrient requirements differ between organisms; for example, hum ...
s in appreciable quantities (table).


Traditional medicine

Longan may be used in traditional Eastern folk medicine. Prior to the 1800s, longan was prevalent in Asia. In ancient Vietnamese medicine, the "eye" of the longan seed was pressed against snakebites to absorb the venom. This was ineffective, but may still be used in the 21st century.


Growth, Harvest, and Distribution


Growth

Longan, alongside its sister fruit litchi, thrive in humid areas or places with high rainfall, and can grow on most types of soil that does not induce issues with water drainage. Ample temperatures are also instrumental in longan growth - while longan can resist small stretches of cool temperatures, they can be damaged or killed in longer stretches of temperatures as high as -2 degrees Celsius. Younger plants tend to be more vulnerable to the cold than those more mature.


Harvest

During harvest, pickers must climb ladders to carefully remove branches of fruit from longan trees. Longan fruit remain fresher if still attached to the branch, so efforts are made to prevent the fruit from detaching too early. Mechanical picking would damage the delicate skin of the fruit, so the preferred method is to harvest by hand. Knives and scissors are the most commonly used tools. Fruit is picked early in the day to minimize water loss and to prevent high heat exposure, which would be damaging. The fruit is then placed into either plastic crates or bamboo baskets and taken to packaging houses, where the fruit undergo a series of checks for quality. The packaging houses are well-ventilated and shaded to prevent further decay. The process of checking and sorting are performed by workers instead of machinery. Any fruit that is split, under-ripe, or decaying is disposed of. The remaining healthy fruit is then prepped and shipped to markets. Many companies add preservatives to canned longan. Regulations control the preserving process. The only known preservative added to canned longan is
sulfur dioxide Sulfur dioxide (IUPAC-recommended spelling) or sulphur dioxide (traditional Commonwealth English) is the chemical compound with the formula . It is a toxic gas responsible for the odor of burnt matches. It is released naturally by volcanic ac ...
, to prevent discoloration. Fresh longan that is shipped worldwide is exposed to sulfur
fumigation Fumigation is a method of pest control or the removal of harmful micro-organisms by completely filling an area with gaseous pesticides—or fumigants—to suffocate or poison the pests within. It is used to control pests in buildings (s ...
. Tests have shown that sulfur residues remain on the fruit skin, branches, and leaves for a few weeks. This violates many countries' limits on fumigation residue, and efforts have been made to reduce this amount.


Distribution

Longan is found commonly in most of Asia, primarily in mainland China, Taiwan, Vietnam and Thailand. China, the main longan-producing country in the world, produced about of longan in 2015–2017, accounting for 70% of the world’s longan production and more than 50% of the world’s longan plots. Vietnam and Thailand produced around , respectively. Like Vietnam, Thailand's economy relies heavily on the cultivation and shipments of longan as well as lychee. This increase in the production of longan reflects recent interest in exotic fruits in other parts of the world. However, the majority of the demand comes from Asian communities in North America, Europe and Australia.


Longan growth enhancement efforts

While longan yields average out to 2 to 5 metric tons per hectare, there have been observed yields of up to 19.5 mt/ha in Israel. When investigating the difference, researchers found that the biggest areas of improvement for lychee yield came in improving crop management practices and the ambience surrounding the fields. Efforts have been made to prolong longan's relatively short shelf life with chemical treatments based on SO2 in 2014. Advancements in selective breeding have allowed scientists to find a strain of longan containing a "high proportion of aborted seeds" at the end of a thirty year breeding program in 2001. Studies in 2015 that aimed to aid longan breeding efforts discovered that -20 degrees Celsius is the optimal temperature for long-term storage of longan pollen, a key ingredient in enabling longan breeding programs.


Diseases

Plant based diseases can affect both longan fruits and their trees, and the severity of these diseases can range from harmless cosmetic damage to rendering to the fruit inedible. The most prevalent disease among longan plants is the ''
Witch's broom Witch's broom or witches' broom is a deformity in a woody plant, typically a tree, where the natural structure of the plant is changed. A dense mass of shoots grows from a single point, with the resulting structure resembling a broom or a ...
'', which can be found in all major longan-producing Asian territories, including China, Thailand, and Vietnam. ''Witch's Broom'' works on deforming longan skin, and at times having the plant prematurely drop their fruit, similar to the '' Phytophthora palmivora''. Another common disease that longan trees can carry is the aptly named ''Longan Decline'', which is largely prevalent in Thailand, with reports finding that it could affect up to 40% of longan trees alone. Affected trees are more vulnerable to common tree pests and algae, and often bear low-quality fruit unworthy of yield. ''Algal spot'' is another plant disease that can affect longan plants and trees. Common among tropical fruits, the disease mainly takes form as red-orange algae that can appear on a fruit-bearing tree's leaves or branches.''Algal spot'' on longan plants, like many other tropical fruits, is caused by Cephaleuros virescens. An oomycete disease that causes
blight Blight refers to a specific symptom affecting plants in response to infection by a pathogenic organism. Description Blight is a rapid and complete chlorosis, browning, then death of plant tissues such as leaves, branches, twigs, or floral orga ...
on leaves and foliage of a plant and affects the related lychee, '' Phytophthora palmivora'', can also appear on both longan plants and fruit, particularly in the Thailand region. When affecting longan, it can create brown spots on the fruit in an erratic fashion, and can also cause longan to drop prematurely from the plant. Early symptoms can also include a dark
necrosis Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. Necrosis is caused by factors external to the cell or tissue, such as infection, or trauma which result in the unregulated dig ...
on the plant itself. Stem-end rot is a disease common amongst litchi and longan, and causes browning and rot on the stem of the fruit. Longan also suffer from various decay-accelerating fungi. An oomycete disease that affects the related lychee - ''
Phytophthora ''Phytophthora'' (from Greek (''phytón''), "plant" and (), "destruction"; "the plant-destroyer") is a genus of plant-damaging oomycetes (water molds), whose member species are capable of causing enormous economic losses on crops worldwide, a ...
litchii'' - also afflicts ''D. longan''.


Gallery

File:Longan (Dimocarpus longan) Tree leaves.jpg, Longan (''Dimocarpus longan'') tree leaves File:Longan (Dimocarpus longan) Tree lower trunk.jpg, Longan tree lower trunk File:Flora_Cochinchinensis,_t._1.jpg, The longan was described in Joao de Loureiro's work, ''Flora Cochinchinensis'', published in 1790 File:Longan (Dimocarpus longan) baby fruits and leaves.jpg, Longan (Dimocarpus longan) baby fruits and leaves


See also

*''
Lansium parasiticum ''Lansium parasiticum'', commonly known as langsat (), lanzones (), or longkong in English; duku in Indonesian or dokong in Terengganu Malay, is a species of tree in the Mahogany family with commercially cultivated edible fruits. The specie ...
'', the langsat or lanzones


References


External links


Longan Production in Asia
from the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
of the United Nations {{Authority control Articles containing video clips Cantonese cuisine Chinese fruit
longan ''Dimocarpus longan'', commonly known as the longan () and dragon's eye, is a tropical tree species that produces edible fruit. It is one of the better-known tropical members of the soapberry family Sapindaceae, to which the lychee and rambu ...
Edible fruits Flora of tropical Asia Fruits originating in East Asia Han dynasty Indomalayan realm flora Near threatened plants Plants described in 1790 Traditional Chinese medicine Trees of China Tropical fruit