The liger is a hybrid offspring of a male

lion The lion (''Panthera leo'') is a large cat of the genus '' Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adu ...

(''Panthera leo'') and a female

tiger The tiger (''Panthera tigris'') is the largest living Felidae, cat species and a member of the genus ''Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily pr ...

(''Panthera tigris''). The liger has parents in the same

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

but of different

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...

. The liger is distinct from the similar hybrid called the

tigon A tigon (), tiglon () (portmanteau of ''tiger'' and ''lion''), or tion () is the hybrid offspring of a male tiger (''Panthera tigris'') and a female lion (''Panthera leo'').

, and is the largest of all known

extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...

felines The Felinae are a subfamily of the family Felidae. This subfamily comprises the small cats having a bony hyoid, because of which they are able to purr but not roar. Other authors have proposed an alternative definition for this subfamily: a ...

. They enjoy swimming, which is a characteristic of tigers, and are very sociable like lions. Notably, ligers typically grow larger than either parent species, unlike tigons.Ligers
messybeast.com Retrieved 4 September 2012.

History

The history of lion-tiger hybrids dates to at least the early 19th century in

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...

. In 1798,

Étienne Geoffroy Saint-Hilaire Étienne Geoffroy Saint-Hilaire (15 April 177219 June 1844) was a French naturalist who established the principle of "unity of composition". He was a colleague of Jean-Baptiste Lamarck and expanded and defended Lamarck's evolutionary theories ...

(1772–1844) made a colour plate of the offspring of a lion and a tiger. The name "liger", a

portmanteau A portmanteau word, or portmanteau (, ) is a blend of wordsG. B. Whittaker made an engraving of liger cubs born in 1824. The parents and their three liger offspring are also depicted with their trainer in a 19th-century painting in the naïve style. Two liger cubs born in 1837 were exhibited to King

William IV William IV (William Henry; 21 August 1765 – 20 June 1837) was King of the United Kingdom of Great Britain and Ireland and King of Hanover from 26 June 1830 until his death in 1837. The third son of George III, William succeeded h ...

and to his successor Queen

Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...

. On 14 December 1900 and on 31 May 1901,

Carl Hagenbeck Carl Hagenbeck (10 June 1844 – 14 April 1913) was a German merchant of wild animals who supplied many European zoos, as well as P. T. Barnum. He created the modern zoo with animal enclosures without bars that were closer to their natural hab ...

wrote to zoologist

James Cossar Ewart James Cossar Ewart FRS FRSE (26 November 1851 – 31 December 1933) was a Scottish zoologist. He performed breeding experiments with horses and zebras which disproved earlier theories of heredity. Life Ewart was born in Penicuik, Midloth ...

with details and photographs of ligers born at the Hagenbeck's Tierpark in Hamburg in 1897. In ''Animal Life and the World of Nature'' (1902–1903), A. H. Bryden described Hagenbeck's "lion-tiger" hybrids:

It has remained for one of the most enterprising collectors and naturalists of our time, Mr. Carl Hagenbeck, not only to breed but to bring successfully to a healthy maturity, specimens of this rare alliance between those two great and formidable Felidae, the lion and tiger. The illustrations will indicate sufficiently how fortunate Mr. Hagenbeck has been in his efforts to produce these hybrids. The oldest and biggest of the animals shown is a hybrid born on the 11th May 1897. This fine beast, now more than five years old, equals and even excels in his proportions a well-grown lion, measuring as he does from nose tip to tail 10 ft 2 inches in length, and standing only three inches less than 4 ft at the shoulder. A good big lion will weigh about 400 lb ..the hybrid in question, weighing as it does no less than 467 lb, is certainly the superior of the most well-grown lions, whether wild-bred or born in a menagerie. This animal shows faint striping and mottling, and, in its characteristics, exhibits strong traces of both its parents. It has a somewhat lion-like head, and the tail is more like that of a lion than of a tiger. On the other hand, it has no trace of mane. It is a huge and very powerful beast.

In 1935, four ligers from two litters were reared in the Zoological Gardens of

Bloemfontein Bloemfontein, ( ; , "fountain of flowers") also known as Bloem, is one of South Africa's three capital cities and the capital of the Free State province. It serves as the country's judicial capital, along with legislative capital Cape To ...

, South Africa. Three of them, a male and two females, were still living in 1953. The male weighed and stood a foot and a half (45 cm) taller than a full grown male lion at the shoulder. Although ligers are more commonly found than tigons today, in ''At Home in the Zoo'' (1961), Gerald Iles wrote "For the record I must say that I have never seen a liger, a hybrid obtained by crossing a lion with a tigress. They seem to be even rarer than tigons."

Appearance

The liger has a faint tiger-like striped pattern upon a lionesque tawny background. In addition, it may inherit rosettes from the lion parent (lion cubs are rosetted and some adults retain faint markings). These markings may be black, dark brown or sandy. The background color may be correspondingly tawny, sandy or golden. In common with tigers, as an example of

countershading Countershading, or Thayer's law, is a method of camouflage in which an animal's coloration is darker on the top or upper side and lighter on the underside of the body. This pattern is found in many species of mammals, reptiles, birds, fish ...

, the underparts are pale. The specific pattern and color depend upon which subspecies the parents were and how the genes interact in the offspring.

White tiger The white tiger or bleached tiger is a leucistic pigmentation variant of the Mainland tiger. It is reported in the wild from time to time in the Indian states of Madhya Pradesh, Assam, West Bengal, Bihar, Odisha, in the Sunderbans region and ...

s have been crossed with lions to produce "white" (actually pale golden) ligers. In theory, white tigers could be crossed with

white lion The white lion is a rare color mutation of the lion, specifically the Southern African lion. White lions in the area of Timbavati are thought to have been indigenous to the Timbavati region of South Africa for centuries, although the earliest ...

s to produce white, very pale or even stripeless ligers. There are no black ligers. Very few melanistic tigers have ever been recorded, most being due to excessive markings (pseudo-melanism or

abundism The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair. Pseudomelanism, also called abundism, is another variant of pi ...

) rather than true melanism; no reports of black lions have ever been substantiated. As blue or Maltese tigers probably no longer exist, gray or blue ligers are exceedingly improbable. It is not impossible for a liger to be white, but it is very rare. The first known white ligers were born in December 2013 at Myrtle Beach Safari in

Myrtle Beach, South Carolina Myrtle Beach is a resort city on the east coast of the United States in Horry County, South Carolina. It is located in the center of a long and continuous stretch of beach known as "The Grand Strand" in the northeastern part of the state. Its ...

to a white male lion and a white female tiger.

Size and growth

The liger is often believed to be the largest cat in the world. Males reach a total length of ,Description of ligers at ''Bestiarium.kryptozoologie.net''
/ref>
/ref> which means that they rival even large male lions and tigers in length.

Vratislav Mazák Vratislav Mazák (; 22 June 1937 – 9 September 1987) was a Czech biologist who specialized in paleoanthropology, mammalogy and taxonomy. He was also a painter, often illustrating his books about animals and men. Born at Kutná Hora, he was a p ...

: ''Der Tiger''. Westarp Wissenschaften; Auflage: 5 (April 2004), unveränd. Aufl. von 1983 Imprinted genes may be a factor contributing to the large size of ligers. These are genes that may or may not be expressed on the parent they are inherited from, and that occasionally play a role in issues of hybrid growth. For example, in some dog breed crosses, genes that are expressed only when maternally-inherited cause the young to grow larger than is typical for either parent breed. This growth is not seen in the paternal breeds, as such genes are normally "counteracted" by genes inherited from the female of the appropriate breed. Other

big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. Despite enormous differences in size, various cat species are quite similar ...

hybrids can reach similar sizes; the litigon, a rare hybrid of a male lion and a female tigon, is roughly the same size as the liger, with a male named Cubanacan (at the

Alipore Zoo The Zoological Garden, Alipore (also informally called the Alipore Zoo or Kolkata Zoo) is India's oldest formally stated zoological park (as opposed to royal and British menageries) and a big tourist attraction in Kolkata, West Bengal. It has b ...

in India) reaching . The extreme rarity of these second-generation hybrids may make it difficult to ascertain whether they are larger or smaller, on average, than the liger. It is sometimes wrongly believed that ligers continue to grow throughout their lives due to hormonal issues. It may be that they simply grow far more during their growing years and take longer to reach their full adult size. Further growth in shoulder height and body length is not seen in ligers over six years old, as in both lions and tigers. Male ligers also have the same levels of

testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteristi ...

on average as an adult male lion, yet are

azoospermic Azoospermia is the medical condition of a man whose semen contains no sperm. It is associated with male infertility, but many forms are amenable to medical treatment. In humans, azoospermia affects about 1% of the male population and may be seen ...

in accordance with Haldane's rule. In addition, female ligers may also attain great size, weighing approximately and reaching long on average, and are often fertile. In contrast,

pumapard A pumapard is a hybrid of a cougar and a leopard. Both male cougar with female leopard and male leopard with female cougar pairings have produced offspring. In general, these hybrids have exhibited a tendency to dwarfism. Characteristics ...

s (hybrids between pumas and

leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, ...

s) tend to exhibit

dwarfism Dwarfism is a condition wherein an organism is exceptionally small, and mostly occurs in the animal kingdom. In humans, it is sometimes defined as an adult height of less than , regardless of sex; the average adult height among people with dw ...

. Ligers are about the same size as the prehistoric '' Smilodon populator'' and

American lion ''Panthera atrox'', better known as the American lion, also called the North American lion, or American cave lion, is an extinct pantherine cat that lived in North America during the Pleistocene epoch and the early Holocene epoch, about 340, ...

Records

Hercules, the largest non-obese liger, is recognised by the ''

Guinness Book of World Records ''Guinness World Records'', known from its inception in 1955 until 1999 as ''The Guinness Book of Records'' and in previous United States editions as ''The Guinness Book of World Records'', is a reference book published annually, listing world ...

'' as the largest living cat on Earth, weighing . Hercules was featured on the ''

Today Show ''Today'' (also called ''The Today Show'' or informally, ''NBC News Today'') is an American news and talk morning television show that airs weekdays from 7:00 a.m. to 11:00 a.m. on NBC. The program debuted on January 14, 1952. It w ...

'', ''

Good Morning America ''Good Morning America'' (often abbreviated as ''GMA'') is an American morning television program that is broadcast on ABC. It debuted on November 3, 1975, and first expanded to weekends with the debut of a Sunday edition on January 3, 1993. ...

'', ''

Anderson Cooper 360 Anderson or Andersson may refer to: Companies * Anderson (Carriage), a company that manufactured automobiles from 1907 to 1910 * Anderson Electric, an early 20th-century electric car * Anderson Greenwood, an industrial manufacturer * Anderson ...

'', ''

Inside Edition ''Inside Edition'' is an American news broadcasting newsmagazine program that is distributed in first-run syndication by CBS Media Ventures. Having premiered on January 9, 1989, it is the longest-running syndicated-newsmagazine program that is no ...

'', and in a ''

Maxim Maxim or Maksim may refer to: Entertainment * ''Maxim'' (magazine), an international men's magazine ** ''Maxim'' (Australia), the Australian edition ** ''Maxim'' (India), the Indian edition *Maxim Radio, ''Maxim'' magazine's radio channel on Sir ...

'' article in 2005, when he was only three years old and already weighed . The Valley of the Kings Animal Sanctuary in

Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...

had a male liger named Nook who weighed over . To compare, the records for the lion and tiger in captivity are under .

Health and longevity

Though ligers typically have a life expectancy of between 13 and 18 years, they are occasionally known to live into their 20s. A ligress named Shasta was born at the Hogle Zoo in

Salt Lake City Salt Lake City (often shortened to Salt Lake and abbreviated as SLC) is the capital and most populous city of Utah, United States. It is the seat of Salt Lake County, the most populous county in Utah. With a population of 200,133 in 2020, th ...

on 14 May 1948 and died in 1972 at age 24. Nook, a liger at a facility in Wisconsin, died in 2007, at 21 years old. Hobbs, a male liger at the

Sierra Safari Zoo The Sierra Safari Zoo is a zoo north of Reno, Nevada, United States. Opening in 1989, it has grown to become the zoo that houses more exotics in the state. It houses a wide variety of exotic animals from around the world, including primates, big ca ...

in Reno, Nevada, lived to almost 15 years of age before succumbing to liver failure, and weighed . NSK-ZOO-liger

''Panthera'' hybrids tend to experience a higher rate of injury and neurological disorder than non-hybrids. Though not universal, ligers and tigons may develop health issues. Organ failure issues have been reported in ligers, in addition to

neurological deficit Neurology (from el, νεῦρον (neûron), "string, nerve" and the suffix -logia, "study of") is the branch of medicine dealing with the diagnosis and treatment of all categories of conditions and disease involving the brain, the spinal c ...

s, sterility, cancer, and

arthritis Arthritis is a term often used to mean any disorder that affects joints. Symptoms generally include joint pain and stiffness. Other symptoms may include redness, warmth, swelling, and decreased range of motion of the affected joints. In som ...

Fertility

The fertility of hybrid big cat females is well documented across a number of different hybrids. This is in accordance with Haldane's rule: in hybrids of animals whose sex is determined by

sex chromosomes A sex chromosome (also referred to as an allosome, heterotypical chromosome, gonosome, heterochromosome, or idiochromosome) is a chromosome that differs from an ordinary autosome in form, size, and behavior. The human sex chromosomes, a typical ...

, if one of the two sexes is absent, rare or sterile, it will be the

heterogametic sex Heterogametic sex (digametic sex) refers to the individuals of a species in which the sex chromosomes are not the same. For example, in humans, males with an X and a Y sex chromosome would be referred to as the heterogametic sex, and females ...

. Male ligers are consequently sterile, while female ligers are not. Ligers and tigons were long thought to be totally sterile. However, in 1943, a fifteen-year-old hybrid between a lion and an island tiger was successfully mated with a lion at the Munich Hellabrunn Zoo. The female cub, though of delicate health, was raised to adulthood. In September 2012, the Russian Novosibirsk Zoo announced the birth of a " liliger", the offspring of a liger mother and a lion father. The cub was named Kiara.

Co-occurrence of parent species

As with the tigon, the liger exists only in captivity. Historically, the

Asiatic lion The Asiatic lion is a population of '' Panthera leo leo'' that today survives in the wild only in India. Since the turn of the 20th century, its range has been restricted to Gir National Park and the surrounding areas in the Indian state of Gujar ...

and the

Bengal tiger The Bengal tiger is a population of the '' Panthera tigris tigris'' subspecies. It ranks among the biggest wild cats alive today. It is considered to belong to the world's charismatic megafauna. The tiger is estimated to have been present i ...

co-occurred in some Asian countries, and there are

legend A legend is a genre of folklore that consists of a narrative featuring human actions, believed or perceived, both by teller and listeners, to have taken place in human history. Narratives in this genre may demonstrate human values, and possess ...

s of male lions mating with tigresses in the wilderness, or of ligers existing there. The two species' ranges are known to overlap in India's Gir National Park, though no ligers were known to live there until the modern era. The range of the

Caspian tiger The Caspian tiger was a ''Panthera tigris tigris'' population native to eastern Turkey, northern Iran, Mesopotamia, the Caucasus around the Caspian Sea, Central Asia to northern Afghanistan, and the Xinjiang region in western China. Until the M ...

has overlapped with that of the lion in places such as northern

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

and eastern

Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...

Zoo policies

Keeping the two species separate has been standard procedure. However, ligers have occurred, and do occur, by accident in captivity. Several AZA zoos are reported to have ligers. The USA holds the greatest population of around 30 ligers. China holds about 20 ligers. There are a few countries worldwide that hold a few, but probably less than 100 exist worldwide. The breeding of ligers and other ''Panthera'' hybrids has come under fire from animal rights activists and organisations, who argue that the health problems experienced by these animals makes their creation immoral. Despite these assertions of immorality, some unlicensed zoos still breed ligers for profit.

References

* ''This article incorporates text fro
messybeast.com
which is released under the

GFDL The GNU Free Documentation License (GNU FDL or simply GFDL) is a copyleft license for free documentation, designed by the Free Software Foundation (FSF) for the GNU Project. It is similar to the GNU General Public License, giving readers th ...

.''

External links

The Richmond palladium and sun-telegram: The Theater , Theatrical Calendar
{{Taxonbar, from=Q182573 Panthera hybrids

History

Appearance

Size and growth

Records

Health and longevity

Fertility

Co-occurrence of parent species

Zoo policies

See also

References

Further reading

External links