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The term kestrel (from french: crécerelle, derivative from , i.e. ratchet) is the
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
given to several
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of predatory birds from the
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Falco''. Kestrels are most easily distinguished by their typical hunting behaviour which is to hover at a height of around over open country and swoop down on ground prey, usually small mammals, lizards or large insects, while other falcons are more adapted for active hunting during flight. Kestrels are notable for usually having mostly brown in their plumage.


Description

Most
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
termed kestrels appear to form a distinct
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
among the falcons, as suggested by comparison of
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
cytochrome ''b''
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called ...
data and morphology. This seems to have diverged from other ''Falco'' around the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fi ...
to
Zanclean The Zanclean is the lowest stage or earliest age on the geologic time scale of the Pliocene. It spans the time between 5.332 ± 0.005 Ma (million years ago) and 3.6 ± 0.005 Ma. It is preceded by the Messinian Age of the Miocene Epoch, a ...
, or about 7–3.5 mya). The most basal "true" kestrels are three species from Africa and its surroundings which lack a malar stripe, and in one case have—like other falcons but unlike other true kestrels—large areas of grey in their wings. Approximately during the Gelasian age (
Late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effe ...
or
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
, around 2.5–2 mya), the main lineage of true kestrels emerged; this contains the species characterized by a malar stripe. This too seems to have
evolved Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variati ...
in Africa and subsequently spread across the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by thei ...
until they reached Australia some time during the Middle Pleistocene, less than one million years ago. This group contains several taxa found on Indian Ocean islands. A group of three predominantly grey species from Africa and Madagascar are usually considered kestrels due to their general shape and habits, but are probably distinct from the true kestrels as outlined above. The American kestrel is the only
New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. ...
species termed "kestrel". The molecular data of Groombridge as well as morphological peculiarities (like grey wings in males and a black ear-spot) and
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
, strongly support the view that this species, among the ''Falco'' falcons, is not a kestrel at all in the
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups ...
sense but perhaps closer to the
hobbies A hobby is considered to be a regular activity that is done for enjoyment, typically during one's leisure time. Hobbies include collecting themed items and objects, engaging in creative and artistic pursuits, playing sports, or pursuing oth ...
. Other recent DNA analysis takes a similar view that the American kestrel is not a true kestrel, but rather than being linked to the hobbies is more likely genetically more closely related to the larger American falcons such as the aplomado falcon, the
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey bac ...
, and
prairie falcon The prairie falcon (''Falco mexicanus'') is a medium-large sized falcon of western North America. It is about the size of a peregrine falcon or a crow, with an average length of 40 cm (16 in), wingspan of approximately 1 meter (40&n ...
.Griffiths, C., 1999, "Phylogeny of the Falconidae Inferred from Molecular and Morphological Data", The Auk 116(1):116-130. Though the species has not been renamed as a result of these genetic analyses, it is thus likely a descendant of archaic larger falcon ancestors evolved through a process of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
to fit a similar small prey niche in the ecosystem as the true kestrels.


Groups

Malar-striped clade or common kestrel group *
Malagasy kestrel The Malagasy kestrel (''Falco newtoni''), also known as the Madagascar kestrel, Malagasy spotted kestrel, Newton's kestrel, Madagascar spotted kestrel, ''katiti'' ( Creole) or ''hitsikitsika'' ( Malagasy), is a small bird of prey of the genus '' ...
, ''Falco newtoni'' * Seychelles kestrel, ''Falco araea'' *
Mauritius kestrel The Mauritius kestrel (''Falco punctatus'') is a bird of prey from the family Falconidae endemic to the forests of Mauritius, where it is restricted to the southwestern plateau's forests, cliffs, and ravines. It is the most distinct of the Indi ...
, ''Falco punctatus'' * Réunion kestrel, ''Falco duboisi'' –
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
(c.1700) * Spotted kestrel, ''Falco moluccensis'', found in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Gui ...
* Nankeen kestrel or Australian kestrel, ''Falco cenchroides'', found in
Australia Australia, officially the Commonwealth of Australia, is a sovereign country comprising the mainland of the Australian continent, the island of Tasmania, and numerous smaller islands. With an area of , Australia is the largest country by ...
and
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torres ...
* Common kestrel or Eurasian kestrel, ''Falco tinnunculus'', found in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, and
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
*
Rock kestrel The rock kestrel (''Falco rupicolus'') is a bird of prey species belonging to the kestrel group of the falcon family Falconidae. It was previously considered a subspecies of the common kestrel (''Falco tinnunculus''). This species occurs in Afri ...
, ''Falco rupicolus'', found in South Africa Basal lineage(s) of true kestrels *
Greater kestrel The greater kestrel (''Falco rupicoloides'') or white-eyed kestrel, is a bird of prey belonging to the falcon family Falconidae. It is one of the largest kestrels and is found in open country in southern and eastern Africa. Description The pl ...
, ''Falco rupicoloides'', found in Eastern to South Africa * Fox kestrel, ''Falco alopex'', found in Equatorial Africa * Lesser kestrel, ''Falco naumanni'', found in southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
, and most of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
except for the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
and equatorial forest areas African grey kestrels (a more distant group) *
Grey kestrel The grey kestrel (''Falco ardosiaceus'') is an African bird of prey belonging to the falcon family Falconidae. Its closest relatives are the banded kestrel and Dickinson's kestrel and the three are sometimes placed in the subgenus ''Dissodectes'' ...
, ''Falco ardosiaceus'', found in Central to Southern Africa * Dickinson's kestrel, ''Falco dickinsoni'', found in Eastern to Southern Africa *
Banded kestrel The banded kestrel (''Falco zoniventris'') is a bird of prey belonging to the falcon family Falconidae. It is endemic to Madagascar and is also known as the Madagascar banded kestrel, barred kestrel or Madagascar barred kestrel. Its closest re ...
, ''Falco zoniventris'', found on
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Afric ...
American kestrel * American kestrel, ''Falco sparverius'', found in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...


References


External links

{{Authority control * Falcons Bird common names Messinian first appearances Extant Miocene first appearances