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Haplogroup O-M122 (also known as Haplogroup O2 (formerly Haplogroup O3)) is an
Eastern Eurasian Y-chromosome haplogroup. The lineage ranges across
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainlan ...
and
East Asia
East Asia is the eastern region of Asia, which is defined in both Geography, geographical and culture, ethno-cultural terms. The modern State (polity), states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. ...
, where it dominates the paternal lineages with extremely high frequencies. It is also significantly present in Central Asia, especially among the
Naiman tribe of Kazakhs.
This lineage is a descendant haplogroup of
haplogroup O-M175.
Origins
Researchers believe that O-M122 first appeared in
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainlan ...
approximately 25,000-30,000 years ago or roughly between 30,000 and 35,000 years ago according to more recent studies (Karmin ''et al.'' 2015, Poznik ''et al.'' 2016, YFull January 4, 2018). In a systematic sampling and genetic screening of an East Asian–specific Y-chromosome haplogroup (O-M122) in 2,332 individuals from diverse East Asian populations, results indicate that the O-M122 lineage is dominant in East Asian populations, with an average frequency of 44.3%. Microsatellite data show that the O-M122 haplotypes are more diverse in Southeast Asia than those in northern East Asia. This suggests a southern origin of the O-M122 mutation to be likely.
It was part of the settlement of East Asia. However, the prehistoric peopling of East Asia by modern humans remains controversial with respect to early population migrations and the place of the O-M122 lineage in these migrations is ambivalent.
Distribution
Although Haplogroup O-M122 appears to be primarily associated with
Chinese people, it also forms a significant component of the Y-chromosome diversity of most modern populations of the East Asian region.
East Asia
Haplogroup O-M122 is found in approximately 53.30% of all modern
Chinese males
(with frequency ranging from 30/101=29.7% among
Pinghua
Pinghua (; Yale: ''Pìhng Wá''; sometimes disambiguated as /) is a pair of Sinitic languages spoken mainly in parts of the Guangxi Zhuang Autonomous Region, with some speakers in Hunan province. Pinghua is a trade language in some areas of Gu ...
-speaking Hans in
Guangxi to 110/148=74.3% among Hans in
Changting,
Fujian
Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its cap ...
), about 40% of
Manchu,
Chinese Mongolian,
Korean
Korean may refer to:
People and culture
* Koreans, ethnic group originating in the Korean Peninsula
* Korean cuisine
* Korean culture
* Korean language
**Korean alphabet, known as Hangul or Chosŏn'gŭl
**Korean dialects and the Jeju language
** ...
, and
Vietnamese
Vietnamese may refer to:
* Something of, from, or related to Vietnam, a country in Southeast Asia
** A citizen of Vietnam. See Demographics of Vietnam.
* Vietnamese people, or Kinh people, a Southeast Asian ethnic group native to Vietnam
** Overse ...
males, about 33.3% to 62% ( and ) of
Filipino
Filipino may refer to:
* Something from or related to the Philippines
** Filipino language, standardized variety of 'Tagalog', the national language and one of the official languages of the Philippines.
** Filipinos, people who are citizens of th ...
males, about 10.5% to 55.6% of
Malaysia
Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
n males, about 10% (4/39
Guide County
Guide County is a county in the east of Qinghai Province, China. It is under the administration of Hainan Tibetan Autonomous Prefecture. In Tibetan it's known as Trika. In 2015 it had a population of 108,800, of which 37.8% Tibetans, and 16.1% o ...
,
Qinghai
Qinghai (; alternately romanized as Tsinghai, Ch'inghai), also known as Kokonor, is a landlocked province in the northwest of the People's Republic of China. It is the fourth largest province of China by area and has the third smallest po ...
) to 45% (22/49
Zhongdian County,
Yunnan
Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
) of
Tibetan males, about 20% (10/50
Shuangbai, northern
Yunnan
Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
) to 44% (8/18
Xishuangbanna, southern
Yunnan
Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
) and of
Yi males, about 25% of
Zhuang and
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
n males, and about 16% and to 20% of
Japanese
Japanese may refer to:
* Something from or related to Japan, an island country in East Asia
* Japanese language, spoken mainly in Japan
* Japanese people, the ethnic group that identifies with Japan through ancestry or culture
** Japanese diaspor ...
males. The distribution of Haplogroup O-M122 stretches far into Asia (approx. 40% of
Dungans
Dungan, Xiao'erjing: ; zh, s=东干族, t=東干族, p=Dōnggān zú, w=Tung1kan1-tsu2, , Xiao'erjing: ; russian: Дунгане, ''Dungane''; ky, Дуңгандар, ''Duñgandar'', دۇنغاندار; kk, Дүңгендер, ''Düñgende ...
, 30% of
Salars, 28% of
Bonan, 24% of
Dongxiang, 18% to 22.8% of
Mongolian citizens in Ulaanbaatar, 11%-15.4% of
Khalkha Mongolian
The Khalkha dialect ( mn, Халх аялгуу / / , ) is a dialect of central Mongolic widely spoken in Mongolia. According to some classifications, the Khalkha dialect includes Southern Mongolian varieties such as ''Shiliin gol'', ''Ulaanch ...
s (Yamamoto et al. 2013) but also as high as 31.1% (Kim et al. 2011), 12% of
Uyghurs
The Uyghurs; ; ; ; zh, s=, t=, p=Wéiwú'ěr, IPA: ( ), alternatively spelled Uighurs, Uygurs or Uigurs, are a Turkic ethnic group originating from and culturally affiliated with the general region of Central and East Asia. The Uyghur ...
, 9% of
Kazakhs but in the Naiman of Kazakhs 65.81%,
6.8% of
Kalmyks (17.1% of
Khoshuud
The Khoshut ( Mongolian: Хошууд,, qoşūd, ; literally "bannermen," from Middle Mongolian ''qosighu'' "flag, banner") are one of the four major tribes of the Oirat people. Originally, Khoshuuds were one of the Khorchin tribes in southeaste ...
, 6.1% of
Dörwöd, 3.3% of
Torguud
The Torghut ( Mongolian: Торгууд, , Torguud), , "Guardsman" are one of the four major subgroups of the Four Oirats. The Torghut nobles traced its descent to the Keraite ruler Tooril; also many Torghuts descended from the Keraites.
Hist ...
, 0% of
Buzawa), 6.2% of
Altaians
The Altai people ( alt, Алтай-кижи, Altai-kizhi), also the Altaians ( alt, Алтайлар, Altailar), are a Turkic ethnic group of indigenous peoples of Siberia mainly living in the Altai Republic, Russia. Several thousand of the Alta ...
, 5.3% of
Kyrgyz,
4.1% of
Uzbeks
The Uzbeks ( uz, , , , ) are a Turkic ethnic group native to the wider Central Asian region, being among the largest Turkic ethnic group in the area. They comprise the majority population of Uzbekistan, next to Kazakh and Karakalpak mino ...
, and 4.0% of
Buryats.
Modern northern Han Chinese Y haplogroups and mtdna match those of ancient northern Han Chinese ancestors 3,000 years ago from the Hengbei archeological site. 89 ancient samples were taken. Y haplogroups O3a, O3a3, M, O2a, Q1a1, and O* were all found in Hengbei samples. Three men who lived in the Neolithic era are the ancestors of 40% of Han Chinese, with their Y haplogroups being subclades of O3a-M324 and they are estimated to have lived 6,800 years ago, 6,500 years ago and 5,400 years ago.
The East Asian O3-M122 Y chromosome Haplogroup is found in large quantities in other Muslims close to the
Hui people like Dongxiang, Bo'an and Salar. The majority of Tibeto-Burmans, Han Chinese, and Ningxia and Liaoning Hui share paternal Y chromosomes of East Asian origin which are unrelated to Middle Easterners and Europeans. In contrast to distant Middle Eastern and Europeans whom the Muslims of China are not related to, East Asians, Han Chinese, and most of the Hui and Dongxiang of Linxia share more genes with each other. This indicates that native East Asian populations converted to Islam and were culturally assimilated to these ethnicities and that Chinese Muslim populations are mostly not descendants of foreigners as claimed by some accounts while only a small minority of them are.
South Asia
Haplogroup O-M122 is restricted among tribal groups of
Northeast India
, native_name_lang = mni
, settlement_type =
, image_skyline =
, image_alt =
, image_caption =
, motto =
, image_map = Northeast india.png
, ...
where it is found at very high frequencies. In
Arunachal Pradesh
Arunachal Pradesh (, ) is a state in Northeastern India. It was formed from the erstwhile North-East Frontier Agency (NEFA) region, and became a state on 20 February 1987. It borders the states of Assam and Nagaland to the south. It shares ...
, it is found at 89% among
Adi Adi or ADI may refer to:
Names and titles
* Adi (mythology), an Asura in Hindu faith who appears in the Matsya Purāṇa
* Adi (name), a given name in Hebrew and a nickname in other languages
* Adi (title), a Fijian title used by females of chi ...
, 82% among
Apatani, and 94% among
Nishi, while the
Naga people
Nagas are various ethnic groups native to northeastern India and northwestern Myanmar. The groups have similar cultures and traditions, and form the majority of population in the Indian states of Nagaland and Manipur and Naga Self-Administere ...
show it at 76% (). In
Meghalaya
Meghalaya (, or , meaning "abode of clouds"; from Sanskrit , "cloud" + , "abode") is a state in northeastern India. Meghalaya was formed on 21 January 1972 by carving out two districts from the state of Assam: (a) the United Khasi Hills and J ...
, 59.2% (42/71) of a sample of
Garos and 31.7% (112/353) of a sample of
Khasis have been found to belong to O-M122. In
Nepal
Nepal (; ne, :ne:नेपाल, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne,
सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in S ...
,
Tamang
The Tamang (; Devanagari: तामाङ; ''tāmāṅ'') are an Tibeto-Burmese ethnic group of Nepal. In Nepal Tamang/Moormi people constitute 5.6% of the Nepalese population at over 1.3 million in 2001, increasing to 1,539,830 as of the 2011 ...
people present a very high frequency of O-M122 (39/45 = 86.7%), while much lower percentages of
Newar (14/66 = 21.2%) and the general population of
Kathmandu
, pushpin_map = Nepal Bagmati Province#Nepal#Asia
, coordinates =
, subdivision_type = Country
, subdivision_name =
, subdivision_type1 = Province
, subdivision_name1 = Bagmati Prov ...
(16/77 = 20.8%) belong to this haplogroup. A study published in 2009 found O-M122 in 52.6% (30/57, including 28 members of O-M117 and two members of O-M134(xM117)) of a sample of
Tharus from a village in
Chitwan District of south-central Nepal, 28.6% (22/77, all O-M117) of a sample of Tharus from another village in Chitwan District, and 18.9% (7/37, all O-M117) of a sample of Tharus from a village in
Morang District
Morang District ( ne, मोरङ जिल्ला ) is located in Province No. 1 in eastern Nepal. It is an Outer Terai district. It borders with Bihar, India to the South, Jhapa to the East, Dhankuta and Panchthar to the North, and Sunsa ...
of southeastern Nepal.
In contrast, the same study found O-M122 in only one individual in a sample of non-Tharu Hindus collected in Chitwan District (1/26 = 3.8% O-M134(xM117)), one
tribal
The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to conflic ...
individual from Andhra Pradesh, India (1/29 = 3.4% O-M117), and one individual in a sample of Hindus from New Delhi, India (1/49 = 2.0% O-M122(xM134)).
Southeast Asia
Among all the populations of East and Southeast Asia, Haplogroup O-M122 is most closely associated with those that speak a
Sinitic
The Sinitic languages (漢語族/汉语族), often synonymous with "Chinese languages", are a group of East Asian analytic languages that constitute the major branch of the Sino-Tibetan language family. It is frequently proposed that there is ...
,
Tibeto-Burman
The Tibeto-Burman languages are the non- Sinitic members of the Sino-Tibetan language family, over 400 of which are spoken throughout the Southeast Asian Massif ("Zomia") as well as parts of East Asia and South Asia. Around 60 million people spea ...
, or
Hmong–Mien language. Haplogroup O-M122 comprises about 50% or more of the total Y-chromosome variation among the populations of each of these language families. The Sinitic and Tibeto-Burman language families are generally believed to be derived from a common
Sino-Tibetan protolanguage, and most linguists place the homeland of the Sino-Tibetan language family somewhere in northern China. The Hmong–Mien languages and cultures, for various archaeological and ethnohistorical reasons, are also generally believed to have derived from a source somewhere north of their current distribution, perhaps in northern or central China. The
Tibetans
The Tibetan people (; ) are an East Asian ethnic group native to Tibet. Their current population is estimated to be around 6.7 million. In addition to the majority living in Tibet Autonomous Region of China, significant numbers of Tibetans liv ...
, however, despite the fact that they speak a language of the Tibeto-Burman language family, have high percentages of the otherwise rare haplogroups
D-M15 and
D3, which are also found at much lower frequencies among the members of some other ethnic groups in East Asia and Central Asia.
Haplogroup O-M122 has been implicated as a diagnostic
genetic marker A genetic marker is a gene or DNA sequence with a known location on a chromosome that can be used to identify individuals or species. It can be described as a variation (which may arise due to mutation or alteration in the genomic loci) that can be ...
of the
Austronesian expansion when it is found in populations of insular
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainlan ...
and
Oceania
Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
. It appears at moderately high frequencies in the
Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
,
Malaysia
Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
, and
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
. Its distribution in Oceania is mostly limited to the traditionally Austronesian culture zones, chiefly
Polynesia
Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
(approx. 25% to 32.5% ). O-M122 is found at generally lower frequencies in coastal and island
Melanesia
Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea.
The region includes the four independent countries of Fiji, Va ...
,
Micronesia
Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...
, and
Taiwanese aboriginal tribes (18% to 27.4% of
Micronesians, and 5% of
Melanesians
Melanesians are the predominant and indigenous inhabitants of Melanesia, in a wide area from Indonesia's New Guinea to as far East as the islands of Vanuatu and Fiji. Most speak either one of the many languages of the Austronesian language f ...
, albeit with reduced frequencies of most subclades.
Haplogroup O-M122
* Y-chromosomes, which are not defined by any identified downstream markers, are actually more common among certain non-
Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
populations than among Han Chinese ones, and the presence of these O-M122* Y-chromosomes among various populations of Central Asia, East Asia, and Oceania is more likely to reflect a very ancient shared ancestry of these populations rather than the result of any historical events. It remains to be seen whether Haplogroup O-M122* Y-chromosomes can be parsed into distinct subclades that display significant geographical or ethnic correlations.
Subclade Distribution
Paragroup O-M122*
Paragroup O2*-M122(xO2a-P197) Y-DNA is quite rare, having been detected only in 2/165 = 1.2% of a sample of
Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
in a pool of samples from mainland China, Taiwan, the Philippines, Vietnam, and Malaysia (n=581), 8/641 = 1.2% of a sample of
Balinese in a pool of samples from western Indonesia (n=960), and 7/350 = 2.0% of a sample of males from
Sumba in a pool of samples from eastern Indonesia (n=957). In the same study, O2*-M122(xO2a-P197) Y-DNA was not observed in a pool of samples from Oceania (n=182).
A paper published by a group of mainly Chinese geneticists in the ''American Journal of Human Genetics'' in 2005 reported the detection of O2*-M122(xO2a-M324) Y-DNA in 1.6% (8/488) of a pool of seven samples of
Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
(3/64 = 4.7% Sichuan, 2/98 = 2.0%
Zibo
Zibo (, ) is a prefecture-level city in central Shandong province, China. It borders the provincial capital Jinan to the west, Tai'an to the southwest, Linyi to the south, Weifang to the east, Dongying to the northeast, and Binzhou to the ...
, Shandong, 1/60 = 1.7% Inner Mongolia, 1/81 = 1.2% Yunnan, 1/86 = 1.2%
Laizhou
Laizhou, alternately romanized as Laichow, is a county-level city in the Prefecture-level city of Yantai, Shandong Province, China. As of 2008, Laizhou had a population of 902,000, out of which 188,000 are urban residents.
Laizhou traditionally ...
, Shandong, 0/39 Guangxi, 0/60 Gansu). O2*-M122(xO2a-M324) Y-DNA also was detected in the following samples of ethnic minorities in China: 5.9% (1/17) Jingpo from Yunnan, 4.3% (2/47) Zhuang from Yunnan, 4.1% (2/49) Lisu from Yunnan, 3.2% (1/31) Wa from Yunnan, 2.6% (1/39) Zhuang from Guangxi, 2.5% (2/80) Bai from Yunnan, 2.4% (1/41) Hani from Yunnan, 2.3% (2/88) Lahu from Yunnan, 2.1% (1/47) Yi from Yunnan, 2.1% (1/48) Miao from Yunnan, 1.5% (2/132) Dai from Yunnan, 1.0% (1/105) Miao from Hunan, and 0.9% (2/225) Yao from Guangxi.
O2*-M122(xO2a-M324) Y-DNA has been found as a singleton (1/156 = 0.6%) in a sample from
Tibet
Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa, Taman ...
. It also has been found as a singleton in a sample of nineteen members of the
Chin people in
Chin State
Chin State (, ) is a state in western Myanmar. The Chin State is bordered by Sagaing Division and Magway Division to the east, Rakhine State to the south, Bangladesh to the south-west, and the Indian states of Mizoram to the west and Ma ...
, Myanmar.
In a paper published in 2011, Korean researchers have reported finding O2*-M122(xO2a-M324) Y-DNA in the following samples: 5.9% (3/51)
Beijing
}
Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 21 ...
Han, 3.1% (2/64)
Filipino
Filipino may refer to:
* Something from or related to the Philippines
** Filipino language, standardized variety of 'Tagalog', the national language and one of the official languages of the Philippines.
** Filipinos, people who are citizens of th ...
, 2.1% (1/48)
Vietnamese
Vietnamese may refer to:
* Something of, from, or related to Vietnam, a country in Southeast Asia
** A citizen of Vietnam. See Demographics of Vietnam.
* Vietnamese people, or Kinh people, a Southeast Asian ethnic group native to Vietnam
** Overse ...
, 1.7% (1/60)
Yunnan
Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
Han, 0.4% (2/506)
Korean
Korean may refer to:
People and culture
* Koreans, ethnic group originating in the Korean Peninsula
* Korean cuisine
* Korean culture
* Korean language
**Korean alphabet, known as Hangul or Chosŏn'gŭl
**Korean dialects and the Jeju language
** ...
.
In 2011, Chinese researchers published a paper reporting their finding of O2*-M122(xO2a-M324) Y-DNA in 3.0% (5/167) of a sample of Han Chinese with origins in East China (defined as consisting of Jiangsu, Zhejiang, Shanghai, and Anhui) and in 1.5% (1/65) of a sample of Han Chinese with origins in Southern China. O2* Y-DNA was not detected in their sample of Han Chinese with origins in Northern China (n=129).
In a paper published in 2012, O2*-M122(xO2a-P200) Y-DNA was found in 12% (3/25) of a sample of
Lao males from
Luang Prabang, Laos. O2* Y-DNA was not detected in this study's samples of Cham from Binh Thuan, Vietnam (n=59), Kinh from Hanoi, Vietnam (n=76), or Thai from northern Thailand (n=17).
Trejaut ''et al.'' (2014) found O2-M122(xO2a-M324) in 6/40 (15.0%)
Siraya in
Kaohsiung, 1/17 (5.9%)
Sulawesi, 1/25 (4.0%)
Paiwan, 2/55 (3.6%)
Fujian
Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its cap ...
Han, 1/30 (3.3%)
Ketagalan, 2/60 (3.3%)
Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
Minnan, 1/34 (2.9%)
Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
Hakka
The Hakka (), sometimes also referred to as Hakka Han, or Hakka Chinese, or Hakkas are a Han Chinese subgroup whose ancestral homes are chiefly in the Hakka-speaking provincial areas of Guangdong, Fujian, Jiangxi, Guangxi, Sichuan, Hunan, Zhej ...
, 1/38 (2.6%)
Siraya in
Hwalien
Hualien City (; Wade-Giles: Hua¹-lien² Shih⁴; Hokkien Pe̍h-ōe-jī, POJ: ''Hoa-lian-chhī'' or ''Hoa-liân-chhī'') is a county-administered city and the county seat of Hualien County, Taiwan. It is located on the east coast of Taiwan on th ...
, 5/258 (1.9%) miscellaneous
Han volunteers in Taiwan, and 1/75 (1.3%) in a sample of the general population of
Thailand
Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
.
Brunelli ''et al.'' (2017) found O2-M122(xO2a-M324) in 5/66 (7.6%)
Tai Yuan, 1/91 (1.1%)
Tai Lue, and 1/205 (0.5%)
Khon Mueang in samples of the people of
Northern Thailand
Northern Thailand, or more specifically Lanna, is geographically characterised by several mountain ranges, which continue from the Shan Hills in bordering Myanmar to Laos, and the river valleys which cut through them. Though like most of Thailan ...
.
O-M324
= O-M121
=
O2a1a1a1a1-M121 is a subclade of O2a1-L127.1, parallel to O2a1b-M164 and O2a1c-JST002611.
In an early survey of Y-DNA variation in present-day human populations of the world, O-M121 was detected only in 5.6% (1/18) of a sample from Cambodia and Laos and in 5.0% (1/20) of a sample from China.
In a large study of 2,332 unrelated male samples collected from 40 populations in East Asia (and especially
Southwest China
Southwest China () is a region in the south of the People's Republic of China.
Geography
Southwest China is a rugged and mountainous region, transitioning between the Tibetan Plateau to the west and the Chinese coastal hills (东南丘陵) and ...
), O-M121/DYS257 Y-DNA was detected only in 7.1% (1/14) of a sample of
Cambodia
Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
ns and in 1.0% (1/98) of a sample of
Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
from
Zibo
Zibo (, ) is a prefecture-level city in central Shandong province, China. It borders the provincial capital Jinan to the west, Tai'an to the southwest, Linyi to the south, Weifang to the east, Dongying to the northeast, and Binzhou to the ...
, Shandong.
In a study published in 2011, O-M121 Y-DNA was found in 1.2% (2/167) of a sample of Han Chinese with origins in East China, defined as consisting of Jiangsu, Anhui, Zhejiang, and Shanghai, and in 0.8% (1/129) of a sample of Han Chinese with origins in Northern China. O-M121 was not detected in this study's sample of Han Chinese with origins in Southern China (n=65).
O-L599 (considered to be phylogenetically equivalent to O-M121
[ISOGG Y-DNA Haplogroup O and its Subclades - 2017](_blank)
/ref>) also has been found in one individual in the 1000 Genomes Project
The 1000 Genomes Project (abbreviated as 1KGP), launched in January 2008, was an international research effort to establish by far the most detailed catalogue of human genetic variation. Scientists planned to sequence the genomes of at least one th ...
sample of Han Chinese from Hunan
Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi ...
, China (''n''=37), one individual in the 1000 Genomes Project sample of Kinh
The Vietnamese people ( vi, người Việt, lit=Viet people) or Kinh people ( vi, người Kinh) are a Southeast Asian ethnic group native to modern-day Northern Vietnam and Southern China (Jing Islands, Dongxing, Guangxi). The native lang ...
from Ho Chi Minh City, Vietnam, one individual in the Human Genome Diversity Project
The Human Genome Diversity Project (HGDP) was started by Stanford University's Morrison Institute in 1990s along with collaboration of scientists around the world. It is the result of many years of work by Luigi Cavalli-Sforza, one of the most ci ...
sample of Tujia, an individual from Singapore, and an individual from the Jakarta metropolitan area
The Jakarta metropolitan area or Greater Jakarta, known locally as Jabodetabek (an acronym of Jakarta– Bogor– Depok–Tangerang– Bekasi), and sometimes extended to Jabodetabekjur (with the acronym extended to include part of Cianjur Regen ...
. According to 23mofang, O-L599 currently accounts for about 0.79% of the male population in China and is concentrated in Fujian, Taiwan, Jiangxi, Anhui, Hubei, Zhejiang and other provinces and cities; it appears to have undergone explosive population growth between about 2600 and 2300 years ago.
= O-M164
=
O2a1b-M164 is a subclade of O2a1-L127.1, parallel to O2a1a1a1a1-M121 and O2a1c-JST002611.
In an early survey of Y-DNA variation in present-day human populations of the world, O-M164 was detected only in 5.6% (1/18) of a sample from Cambodia and Laos.
In a large study of 2,332 unrelated male samples collected from 40 populations in East Asia (and especially Southwest China), O2a1b-M164 Y-DNA was detected only in 7.1% (1/14) of a sample of Cambodia
Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
ns.
According to 23mofang, O-M164 is a recent branch (TMRCA 2120 years) downstream of O2a1c-JST002611 rather than parallel to it. Out of fourteen members total, six are from Guangdong, five are from Fujian, one is from Nantong
Nantong (; alternate names: Nan-t'ung, Nantung, Tongzhou, or Tungchow; Qihai dialect: ) is a prefecture-level city in southeastern Jiangsu province of China, province, China. Located on the northern bank of the Yangtze River, near the river mouth. ...
, one is from Wenzhou
Wenzhou (pronounced ; Wenzhounese: Yuziou �y33–11 tɕiɤu33–32 ), historically known as Wenchow is a prefecture-level city in southeastern Zhejiang province in the People's Republic of China. Wenzhou is located at the extreme south east ...
, and one is from Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
.[Phylogenetic tree of human Y-DNA at 23mofang](_blank)
/ref>
= O-JST002611
=
Haplogroup O2a1c-JST002611 is derived from O2-M122 via O2a-M324/P93/P197/P199/P200 and O2a1-L127.1/L465/L467. O2a1c-JST002611 is the most commonly observed type of O2a1 Y-DNA, and, more generally, represents the majority of extant O2-M122 Y-DNA that does not belong to the expansive subclade O2a2-P201.
Haplogroup O2a1c-JST002611 was first identified in 3.8% (10/263) of a sample of Japanese
Japanese may refer to:
* Something from or related to Japan, an island country in East Asia
* Japanese language, spoken mainly in Japan
* Japanese people, the ethnic group that identifies with Japan through ancestry or culture
** Japanese diaspor ...
(Nonaka ''et al.'' 2007). It also has been found in 3.5% (2/57) of the JPT (Japanese in Tokyo, Japan) sample of the 1000 Genomes Project
The 1000 Genomes Project (abbreviated as 1KGP), launched in January 2008, was an international research effort to establish by far the most detailed catalogue of human genetic variation. Scientists planned to sequence the genomes of at least one th ...
, including one member of the rare and deeply divergent paragroup O2a1c1-F18*(xO2a1c1a1-F117, O2a1c1a2-F449).[YFull](_blank)
Haplogroup YTree v6.01 at 4 January 2018 Subsequently, this haplogroup has been found with higher frequency in some samples taken in and around China, including 12/58 = 20.7% Miao (China), 10/70 = 14.3% Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
, 18/165 = 10.9% Han (China & Taiwan), 4/49 = 8.2% Tujia (China). O-002611 also has been found in a singleton from the Philippines (1/48 = 2.1%), but it has not been detected in samples from Malaysia (0/32), Taiwanese Aboriginals (0/48), She from China (0/51), Yao from China (0/60), Oceania (0/182), eastern Indonesia (0/957), or western Indonesia (0/960). Haplogroup O2a1c‐JST002611 is prevalent in different
ethnic groups in China and Southeast Asia, including Vietnam (14.29%), Sichuan of southwestern China (Han, 14.60%; Tibetan in Xinlong County, 15.22%), Jilin of northeastern China (Korean, 9.36%), Inner Mongolia (Mongolian, 6.58%), and Gansu of northwestern China (Baima, 7.35%; Han, 11.30%). Y-DNA belonging to haplogroup O-JST002611 has been observed in 10.6% (61/573) of a sample collected in Seoul and 8.3% (11/133) of a sample collected in Daejeon, South Korea.
According to 23mofang, haplogroup O-IMS-JST002611 currently accounts for approximately 14.69% of the entire male population of China, and its TMRCA is estimated to be 12,770 years.
O-P201
O2a2-JST021354/P201 is a subclade of O2a that includes the most common types of O2-M122 Y-DNA. This clade includes the major subclades O2a2b1-M134 (subclade of O-P164) and O2a2a1a2-M7, which exhibit expansive distributions centered on China, as well as an assortment of Y-chromosomes that have not yet been assigned to any subclade.
O2a2-P201(xO2a2a1a2-M7, O2a2b1-M134) Y-DNA has been detected with high frequency in many samples of Austronesian-speaking populations, in particular some samples of Batak Toba from Sumatra (21/38 = 55.3%), Tongans
Tongans, a Polynesians, Polynesian group, represent more than 98% of the inhabitants of Tonga. The rest are European (the majority are British people, British), mixed European, and other Pacific Islanders. There also are several hundred Chinese i ...
(5/12 = 41.7%), and Filipinos
Filipinos ( tl, Mga Pilipino) are the people who are citizens of or native to the Philippines. The majority of Filipinos today come from various Austronesian ethnolinguistic groups, all typically speaking either Filipino, English and/or othe ...
(12/48 = 25.0%). Outside of Austronesia, O2a2-P201(xO2a2a1a2-M7, O2a2b1-M134) Y-DNA has been observed in samples of Tujia (7/49 = 14.3%), Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
(14/165 = 8.5%), Japanese
Japanese may refer to:
* Something from or related to Japan, an island country in East Asia
* Japanese language, spoken mainly in Japan
* Japanese people, the ethnic group that identifies with Japan through ancestry or culture
** Japanese diaspor ...
(11/263 = 4.2%), Miao (1/58 = 1.7%), and Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
(1/70 = 1.4%) ( and ).
O-M159
O2a2a1a1a-M159 is a subclade of O2a2-P201. In an early survey of Y-DNA variation in present-day human populations of the world, O-M159 was detected only in 5.0% (1/20) of a sample from China.
Unlike its phylogenetic siblings, O-M7 and O-M134, O-M159 is very rare, having been found only in 2.9% (1/35) of a sample of Han males from Meixian, Guangdong in a study of 988 males from East Asia.
In a study published in 2011, O-M159 was detected in 1.5% (1/65) of a sample of Han Chinese with origins in Southern China. O-M159 was not detected in the same study's samples of Han Chinese with origins in East China (n=167) or Northern China (n=129).
Trejaut ''et al.'' (2014) found O-M159 in 5.0% (3/60) Minnan in Taiwan, 4.2% (1/24) Hanoi, Vietnam, 3.88% (10/258) miscellaneous Han volunteers in Taiwan, 3.6% (2/55) Han in Fujian, 3.24% (12/370) Plains Aborigines in Taiwan (mostly assimilated to Han Chinese), 1.04% (2/192) Western Indonesia (1/25 Kalimantan, 1/26 Sumatra), and 0.68% (1/146) Philippines (1/55 South Luzon).
Kutanan ''et al.'' (2019) found O-M159 in 1.6% (2/129) of their samples of Thai people
Thai people ( th, ชาวไทย; '' endonym''), Central Thai people ( th, คนภาคกลาง, sou, คนใต้, ตามโพร; ''exonym and also domestically'') or Siamese ( th, ชาวสยาม; ''historical exonym an ...
from Central Thailand.
According to 23mofang, the TMRCA of haplogroup O-M159 is estimated to be 8,020 years. It is currently distributed mainly in southern China, accounting for about 0.78% of the total male population of China.
O-M7
Haplogroup O2a2a1a2-M7 Y-DNA has been detected with high frequency in some samples of populations who speak Hmong-Mien languages, Katuic languages, or Bahnaric languages, scattered through some mostly mountainous areas of southern China, Laos, and Vietnam
Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
.
O-M7 has been noted for having a widespread but uneven distribution among populations that speak Hmong-Mien languages, such as She (29/51 = 56.9% She, 10/34 = 29.4% She, 14/56 = 25.0% Northern She from Zhejiang), Miao (21/58 = 36.2% Miao from China, 17/51 = 33.3% Hmong Daw from northern Laos, 6/49 = 12.2% Yunnan Miao, 2/49 = 4.1% Guizhou Miao, 4/100 = 4.0% Hunan Miao), and Yao (18/35 = 51.4% Yao from Liannan, Guangdong, 29/60 = 48.3% Yao from Guangxi, 12/35 = 34.3% Yao from Bama, Guangxi, 12/37 = 32.4% Zaomin from Guangdong, 5/36 = 13.9% Bunu from Guangxi, 1/11 = 9.1% Top-Board Mien, 3/41 = 7.3% Native Mien, 2/31 = 6.5% Southern Mien from Guangxi, 1/19 = 5.3% Flowery-Headed Mien from Guangxi, 1/20 = 5.0% Mountain Straggler Mien from Hunan, 1/28 = 3.6% Blue Kimmun from Guangxi, 1/31 = 3.2% Pahng from Guangxi, 1/47 = 2.1% Western Mien from Yunnan, 0/11 Thin Board Mien, 0/31 Lowland Yao from Guangxi, 0/32 Mountain Kimmun from Yunnan, 0/33 Northern Mien, and 0/41 Lowland Kimmun from Guangxi).
Cai ''et al.'' 2010 have reported finding high frequencies of O-M7 in their samples of Katuic (17/35 = 48.6% Ngeq, 10/45 = 22.2% Katu
KATU (channel 2) is a television station in Portland, Oregon, United States, affiliated with ABC. It is owned by Sinclair Broadcast Group alongside La Grande–licensed Univision affiliate KUNP (channel 16). Both stations share studios on N ...
, 6/37 = 16.2% Kataang, 3/34 = 8.8% Inh (Ir), 4/50 = 8.0% So, 1/39 = 2.6% Suy) and Bahnaric (15/32 = 46.9% Jeh
Jahi is the Avestan language name of Zoroastrianism's demoness of "lasciviousness." As a hypostatic entity, Jahi is variously interpreted as "hussy," "rake," "libertine," "courtesan" and "one who leads a licentious life." Her standard epithet i ...
, 17/50 = 34.0% Oy, 8/32 = 25.0% Brau, 8/35 = 22.9% Talieng, 4/30 = 13.3% Alak, 6/50 = 12.0% Laven) peoples from southern Laos. However, O-M7 has been found only with low frequency in samples of linguistically related Khmuic populations from northern Laos (1/50 = 2.0% Mal, 1/51 = 2.0% Khmu, 0/28 Bit, 0/29 Xinhmul), Vietic peoples from Vietnam and central Laos (8/76 = 10.5% Kinh from Hanoi, Vietnam,[He J-D, Peng M-S, Quang HH, Dang KP, Trieu AV, ''et al.'' (2012), "Patrilineal Perspective on the Austronesian Diffusion in Mainland Southeast Asia." ''PLoS ONE'' 7(5): e36437. doi:10.1371/journal.pone.0036437] 4/50 = 8.0% Kinh
The Vietnamese people ( vi, người Việt, lit=Viet people) or Kinh people ( vi, người Kinh) are a Southeast Asian ethnic group native to modern-day Northern Vietnam and Southern China (Jing Islands, Dongxing, Guangxi). The native lang ...
from northern Vietnam,[Enrico Macholdt, Leonardo Arias, Nguyen Thuy Duong, ''et al.'', "The paternal and maternal genetic history of Vietnamese populations." ''European Journal of Human Genetics'' (2020) 28:636–645. https://doi.org/10.1038/s41431-019-0557-4] 2/28 = 7.1% Bo,[Cai X, Qin Z, Wen B, Xu S, Wang Y, ''et al.'' (2011), "Human Migration through Bottlenecks from Southeast Asia into East Asia during Last Glacial Maximum Revealed by Y Chromosomes." ''PLoS ONE'' 6(8): e24282. doi:10.1371/journal.pone.0024282] 4/70 = 5.7% Vietnamese,[Tatiana M. Karafet, Brian Hallmark, Murray P. Cox, ''et al.'' (2010), "Major East–West Division Underlies Y Chromosome Stratification across Indonesia." ''Mol. Biol. Evol.'' 27(8):1833–1844. doi:10.1093/molbev/msq063] 0/12 Muong, 0/15 Kinh, 0/38 Aheu), Palaungic
The nearly thirty Palaungic or Palaung–Wa languages form a branch of the Austroasiatic languages.
Phonological developments
Most of the Palaungic languages lost the contrastive voicing of the ancestral Austroasiatic consonants, with the disti ...
peoples from northwestern Laos and southwestern Yunnan (2/35 = 5.7% Lamet, 0/29 Ava, 0/52 Blang), and Pakanic peoples from southeastern Yunnan and northwestern Guangxi (0/30 Palyu, 0/32 Bugan).
Haplogroup O-M7 has been found with notable frequency in some samples of Austronesian populations from the central part of the Malay Archipelago (17/86 = 19.8% Indonesians from Borneo, 4/32 = 12.5% Malaysia, 7/61 = 11.5% Java (mostly sampled in Dieng), 6/56 = 10.7% Sumatra,[Manfred Kayser, Ying Choi, Mannis van Oven, ''et al.'' (2008), "The Impact of the Austronesian Expansion: Evidence from mtDNA and Y Chromosome Diversity in the Admiralty Islands of Melanesia." ''Mol. Biol. Evol.'' 25(7):1362–1374. doi:10.1093/molbev/msn078] 4/53 = 7.5% Java, 1/17 = 5.9% Malaysia), but the frequency of this haplogroup appears to drop off very quickly toward the east (1/48 = 2.1% Philippines, 5/641 = 0.8% Balinese, 0/9 Timor
Timor is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is divided between the sovereign states of East Timor on the eastern part and Indonesia on the western part. The Indonesian part, also ...
, 0/28 Alor, 0/30 Moluccas, 0/31 Nusa Tenggaras, 0/33 Moluccas
The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, 0/37 Philippines, 0/40 Borneo, 0/48 Taiwanese Aboriginals, 0/54 Mandar from Sulawesi, 0/92 Lembata
Lembata is an island in the Lesser Sunda Islands, also known as Lomblen island; it is the largest island of the Solor Archipelago, in the Lesser Sunda Islands, Indonesia. It forms a separate regency of the province of Nusa Tenggara Timur. The ...
, 0/350 Sumba, 0/394 Flores) and toward the west (0/38 Batak Toba from Sumatra, 0/60 Nias, 0/74 Mentawai). O-M7 has been found in 14.8% (4/27) of a sample of Giarai from southern Vietnam, 8.3% (2/24) of a sample of Ede from southern Vietnam, and 5.1% (3/59) of a sample of Cham
Cham or CHAM may refer to:
Ethnicities and languages
*Chams, people in Vietnam and Cambodia
**Cham language, the language of the Cham people
***Cham script
*** Cham (Unicode block), a block of Unicode characters of the Cham script
*Cham Albania ...
from Binh Thuan, Vietnam. These Chamic-speaking peoples inhabit southern Vietnam and eastern Cambodia, but their languages are related to those of the Acehnese and Malays. O-M7 also has been found in 21.1% (8/38) of a small set of samples of highlanders of northern Luzon
Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as ...
(including 1/1 Ifugao, 1/2 Ibaloi
The Ibaloi (also spelled Ibaloy; Ibaloi: ''ivadoy'', ) are an indigenous ethnic group found in Benguet Province of the northern Philippines.
''Ibaloi'' is derived from ''i-'', a prefix signifying "pertaining to" and ''badoy'' or house, together ...
, 4/12 Kalangoya, and 2/6 Kankanaey).[Frederick Delfin, Jazelyn M Salvador, Gayvelline C Calacal, ''et al.'', "The Y-chromosome landscape of the Philippines: extensive heterogeneity and varying genetic affinities of Negrito and non-Negrito groups." ''European Journal of Human Genetics'' (2011) 19, 224–230; doi:10.1038/ejhg.2010.162; published online 29 September 2010.]
In the northern fringes of its distribution, O-M7 has been found in samples of Oroqen (2/31 = 6.5%), Tujia from Hunan (3/49 = 6.1%), Qiang (2/33 = 6.1%), Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
(2/32 = 6.3% Han from Yili, Xinjiang, 4/66 = 6.1% Han from Huize
Huize County (, old name: ''Dongchuan 東川'') is a county-level city, under the jurisdiction of Qujing City, Yunnan province, People's Republic of China.
Demography
The city has grown considerably over the past 20 years. It has 871,200 inhabit ...
, Yunnan, 2/35 = 5.7% Han from Meixian, Guangdong, 1/18 = 5.6% Han from Wuhan, Hubei, 6/148 = 4.1% Han from Changting, Fujian, 20/530 = 3.8% Han Chinese from Chongming Island
Chongming, formerly known as Chungming, is an alluvial island at the mouth of the Yangtze River in eastern China covering as of 2010. Together with the islands Changxing and Hengsha, it forms Chongming District, the northernmost area of the ...
,[Zhang, X.; Tang, Z.; Wang, B.; Zhou, X.; Zhou, L.; Zhang, G.; Tian, J.; Zhao, Y.; Yao, Z.; Tian, L.; ''et al.'' "Forensic Analysis and Genetic Structure Construction of Chinese Chongming Island Han Based on Y Chromosome STRs and SNPs." ''Genes'' 2022, 13, 1363. https://doi.org/10.3390/genes13081363] 2/63 = 3.2% Han from Weicheng, Sichuan, 18/689 = 2.6% Han Chinese from Pudong
Pudong is a district of Shanghai located east of the Huangpu, the river which flows through central Shanghai. The name ''Pudong'' was originally applied to the Huangpu's east bank, directly across from the west bank or Puxi, the historic cit ...
,[Manchus
The Manchus (; ) are a Tungusic East Asian ethnic group native to Manchuria in Northeast Asia. They are an officially recognized ethnic minority in China and the people from whom Manchuria derives its name. The Later Jin (1616–1636) and ...]
(1/50 = 2.0% Manchu from
Liaoning[Karafet TM, Osipova LP, Savina OV, Hallmark B, Hammer MF. "Siberian genetic diversity reveals complex origins of the Samoyedic-speaking populations." ''Am J Hum Biol.'' 2018;e23194. https://doi.org/10.1002/ajhb.23194]), and
Koreans
Koreans ( South Korean: , , North Korean: , ; see names of Korea) are an East Asian ethnic group native to the Korean Peninsula.
Koreans mainly live in the two Korean nation states: North Korea and South Korea (collectively and simply re ...
(2/133 = 1.5% Daejeon,
1/300 = 0.3% unrelated Korean males obtained from the National Biobank of Korea,
[Myung Jin Park, Hwan Young Lee, Na Young Kim, ''et al.'' (2013), "Y-SNP miniplexes for East Asian Y-chromosomal haplogroup determination in degraded DNA." ''Forensic Science International: Genetics'' 7, 75–81. http://dx.doi.org/10.1016/j.fsigen.2012.06.014] 1/573 = 0.2% Seoul
).
According to 23mofang, O-M7 has a TMRCA of approximately 13,700 years and is currently relatively common among many ethnic groups in Sichuan and Yunnan, as well as among the Zhuang, Austroasiatic, and Austronesian groups. O-M7 now accounts for about 2.19% of the total male population in China.
O-M134
= O-M134*
=
Paragroup O-M134(xM117) has been found with very high frequency in some samples of
Kim Mun people, a subgroup of the
Yao people
The Yao people (its majority branch is also known as Mien; ; vi, người Dao) is a government classification for various minorities in China and Vietnam. They are one of the 55 officially recognised ethnic minorities in China and reside in t ...
of southern China (16/32 = 50.0% Mountain Kimmun from southern Yunnan, 11/28 = 39.3% Blue Kimmun from western Guangxi). However, this paragroup has been detected in only 3/41 = 7.3% of a sample of Lowland Kimmun from eastern Guangxi. This paragroup also has been found with high frequency in some Kazakh samples, especially the Naiman tribe (102/155 = 65.81%)( and ) Dulik hypothesizes that O-M134 in Kazakhs was due to a later expansion due to its much more recent TMRCA time.
The general outline of the distribution of O-M134(xM117) among modern populations is different as that of the related clade O-M117. In particular, O-M134(xM117) occurs with only low frequency or is nonexistent among most
Tibeto-Burman
The Tibeto-Burman languages are the non- Sinitic members of the Sino-Tibetan language family, over 400 of which are spoken throughout the Southeast Asian Massif ("Zomia") as well as parts of East Asia and South Asia. Around 60 million people spea ...
-speaking populations of Southwest China, Northeast India, and Nepal, who exhibit extremely high frequencies of O-M117. This paragroup also occurs with very low frequency or is non-existent among most Mon-Khmer population of Laos, who exhibit much higher frequencies of O-M117. In Han Chinese, the paragroup is found in approximately the same percentage as O-M117, but has a higher distribution in northern Han Chinese than Southern Han Chinese.
According to 23mofang, the TMRCA of O-M134 is estimated to be 17,900 years, and O-M134(xM117) can be divided into two subsets: O-F122 (TMRCA 17,380 years), which is subsumed alongside O-M117 in an O-F450 clade (TMRCA 17,800 years), and O-MF59333 (TMRCA 13,130 years, currently distributed mainly in southern China and accounting for the Y-DNA of approximately 0.06% of the total male population of China), which is derived from O-M134 but basal to O-F450. O-F122 in turn is divided into O-MF38 (TMRCA 4,560 years, currently distributed mainly in northern China and accounting for the Y-DNA of approximately 0.03% of the total male population of China) and O-F114 (TMRCA 15,310 years, accounts for the Y-DNA of approximately 11.39% of the total male population of China).
The O-F46 (TMRCA 9,190 years) subclade of O-F114 by itself accounts for the Y-DNA of approximately 10.17% of the total male population of present-day China.
In a study of Koreans from
Seoul
Seoul (; ; ), officially known as the Seoul Special City, is the capital and largest metropolis of South Korea.Before 1972, Seoul was the ''de jure'' capital of the Democratic People's Republic of Korea (North Korea) as stated iArticle 103 of ...
(''n''=573) and
Daejeon
Daejeon () is South Korea's fifth-largest metropolis, with a population of 1.5 million as of 2019. Located in the central-west region of South Korea alongside forested hills and the Geum River, the city is known both for its technology an ...
(''n''=133), haplogroup O-M134(xM117), all members of which have been found to belong to O-F444
(phylogenetically equivalent to O-F114
), has been found in 9.42% of the sample from Seoul and 10.53% of the sample from Daejeon.
In a study of Japanese (''n''=263), haplogroup O-M134(xM117) has been observed in nine individuals, or 3.4% of the entire sample set.
The Japanese members of O-M134(xM117) in this study have originated from
Shizuoka (3/12 = 25%),
Tokyo
Tokyo (; ja, 東京, , ), officially the Tokyo Metropolis ( ja, 東京都, label=none, ), is the capital and List of cities in Japan, largest city of Japan. Formerly known as Edo, its metropolitan area () is the most populous in the world, ...
(2/52 = 3.8%),
Toyama (1/3),
Ishikawa (1/4),
Tochigi (1/5), and
Ibaraki (1/5), respectively.
[Sae Naitoh, Iku Kasahara-Nonaka, Kiyoshi Minaguchi, and Phrabhakaran Nambiar, "Assignment of Y-chromosomal SNPs found in Japanese population to Y-chromosomal haplogroup tree." ''Journal of Human Genetics'' (2013) 58, 195–201; doi:10.1038/jhg.2012.159; published online 7 February 2013.]
= O-M117
=
Haplogroup O2a2b1a1-M117 (also defined by the phylogenetically equivalent mutation Page23) is a subclade of O2a2b1-M134 that occurs frequently in
China and in neighboring countries, especially among
Tibeto-Burman
The Tibeto-Burman languages are the non- Sinitic members of the Sino-Tibetan language family, over 400 of which are spoken throughout the Southeast Asian Massif ("Zomia") as well as parts of East Asia and South Asia. Around 60 million people spea ...
-speaking peoples.
O-M117 has been detected in samples of
Tamang
The Tamang (; Devanagari: तामाङ; ''tāmāṅ'') are an Tibeto-Burmese ethnic group of Nepal. In Nepal Tamang/Moormi people constitute 5.6% of the Nepalese population at over 1.3 million in 2001, increasing to 1,539,830 as of the 2011 ...
(38/45 = 84.4%),
Tibetans
The Tibetan people (; ) are an East Asian ethnic group native to Tibet. Their current population is estimated to be around 6.7 million. In addition to the majority living in Tibet Autonomous Region of China, significant numbers of Tibetans liv ...
(45/156 = 28.8% or 13/35 = 37.1%),
Tharus (57/171 = 33.3%), Han Taiwanese (40/183 = 21.9%),
Newars (14/66 = 21.2%), the general population of
Kathmandu
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, coordinates =
, subdivision_type = Country
, subdivision_name =
, subdivision_type1 = Province
, subdivision_name1 = Bagmati Prov ...
, Nepal (13/77 = 16.9%),
Han Chinese
The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctiv ...
(5/34 = 14.7% Chengdu, 5/35 = 14.3% Harbin, 4/35 = 11.4% Meixian, 3/30 = 10.0% Lanzhou, 2/32 = 6.3% Yili),
Tungusic peoples
Tungusic peoples are an ethno-linguistic group formed by the speakers of Tungusic languages (or Manchu–Tungus languages). They are native to Siberia and Northeast Asia.
The Tungusic phylum is divided into two main branches, northern (Evenic or ...
from the PRC (7/45 = 15.6% Hezhe, 4/26 = 15.4% Ewenki, 5/35 = 14.3% Manchu, 2/41 = 4.9% Xibe, 1/31 = 3.2% Oroqen),
Koreans
Koreans ( South Korean: , , North Korean: , ; see names of Korea) are an East Asian ethnic group native to the Korean Peninsula.
Koreans mainly live in the two Korean nation states: North Korea and South Korea (collectively and simply re ...
(4/25 = 16.0% Koreans from the PRC, 5/43 = 11.6% Koreans from South Korea),
Mongols
The Mongols ( mn, Монголчууд, , , ; ; russian: Монголы) are an East Asian ethnic group native to Mongolia, Inner Mongolia in China and the Buryatia Republic of the Russian Federation. The Mongols are the principal membe ...
(5/45 = 11.1% Inner Mongolian, 3/39 = 7.7% Daur, 3/65 = 4.6% Outer Mongolian), and
Uyghurs
The Uyghurs; ; ; ; zh, s=, t=, p=Wéiwú'ěr, IPA: ( ), alternatively spelled Uighurs, Uygurs or Uigurs, are a Turkic ethnic group originating from and culturally affiliated with the general region of Central and East Asia. The Uyghur ...
(2/39 = 5.1%
Yili, 1/31 = 3.2%
Urumqi) (, , and ).
Like O-M7, O-M117 has been found with greatly varying frequency in many samples of
Hmong-Mien-speaking peoples, such as Mienic peoples (7/20 = 35.0% Mountain Straggler Mien, 9/28 = 32.1% Blue Kimmun, 6/19 = 31.6% Flower Head Mien, 3/11 = 27.3% Top Board Mien, 3/11 = 27.3% Thin Board Mien, 11/47 = 23.4% Western Mien, 6/33 = 18.2% Northern Mien, 5/31 = 16.1% Lowland Yao, 5/35 = 14.3% Yao from Liannan, Guangdong, 5/37 = 13.5% Zaomin, 5/41 = 12.2% Lowland Kimmun, 3/41 = 7.3% Native Mien, 2/31 = 6.5% Southern Mien, 2/32 = 6.3% Mountain Kimmun, but 0/35 Yao from Bama, Guangxi), She (6/34 = 17.6% She, 4/56 = 7.1% Northern She), and Hmongic peoples (9/100 = 9.0% Miao from Hunan, 4/51 = 7.8% Hmong Daw from northern Laos, 3/49 = 6.1% Miao from Yunnan, 1/49 = 2.0% Miao from Guizhou, but 0/36 Bunu from Guangxi) ( and ).
In a study published by Chinese researchers in the year 2006, O-M117 has been found with high frequency (8/47 = 17.0%) in a sample of Japanese that should be from
Kagawa Prefecture according to the geographical coordinates (134.0°E, 34.2°N) that have been provided. However, in a study published by Japanese researchers in the year 2007, the same haplogroup has been found with much lower frequency (11/263 = 4.2%) in a larger sample of Japanese from various regions of Japan. More precisely, the Japanese members of O-M117 in this study's sample set have originated from Tokyo (4/52), Chiba (2/44), Gifu (1/2), Yamanashi (1/2), Hiroshima (1/3), Aichi (1/6), and Shizuoka (1/12).
In
Meghalaya
Meghalaya (, or , meaning "abode of clouds"; from Sanskrit , "cloud" + , "abode") is a state in northeastern India. Meghalaya was formed on 21 January 1972 by carving out two districts from the state of Assam: (a) the United Khasi Hills and J ...
, a predominantly tribal state of Northeast India, O-M133 has been found in 19.7% (14/71) of a sample of the
Tibeto-Burman
The Tibeto-Burman languages are the non- Sinitic members of the Sino-Tibetan language family, over 400 of which are spoken throughout the Southeast Asian Massif ("Zomia") as well as parts of East Asia and South Asia. Around 60 million people spea ...
-speaking
Garos, but in only 6.2% (22/353, ranging from 0/32 Bhoi to 6/44 = 13.6%
Pnar) of a pool of eight samples of the neighboring
Khasian-speaking tribes.
O-M300
O-M333
Phylogenetics
Phylogenetic History
Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium (YCC). They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.
Original Research Publications
The following research teams per their publications were represented in the creation of the YCC Tree.
Phylogenetic Trees
This phylogenetic tree of haplogroup O subclades is based on the YCC 2008 tree and subsequent published research.
*O-M122 (M122, P198)
**O-P93 (M324, P93, P197, P198, P199, P200)
***O-M121 (M121, P27.2)
***O-M164 (M164)
***O-P201 (P201/021354)
***O-002611 (002611)
***O-M300 (M300)
***O-M333 (M333)
See also
Genetics
Y-DNA O Subclades
Y-DNA Backbone Tree
References
Citations
Sources
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Sources for conversion tables
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Further reading
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External links
Spread of Haplogroup O-M122 from
The Genographic Project, ''
National Geographic''
China DNAinterest group at
Facebook
Facebook is an online social media and social networking service owned by American company Meta Platforms. Founded in 2004 by Mark Zuckerberg with fellow Harvard College students and roommates Eduardo Saverin, Andrew McCollum, Dustin Mosk ...
China DNA Project Websiteat
Family Tree DNA
FamilyTreeDNA is a division of Gene by Gene, a commercial genetic testing company based in Houston, Texas. FamilyTreeDNA offers analysis of autosomal DNA, Y-DNA, and mitochondrial DNA to individuals for genealogical purpose. With a database of ...
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O-M122