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Guava () is a common tropical fruit cultivated in many tropical and subtropical regions. The common guava '' Psidium guajava'' (lemon guava, apple guava) is a small tree in the myrtle family (
Myrtaceae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All speci ...
), native to Mexico, Central America, the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
and
northern South America The World Geographical Scheme for Recording Plant Distributions (WGSRPD) is a biogeographical system developed by the international Biodiversity Information Standards (TDWG) organization, formerly the International Working Group on Taxonomic Da ...
. The name guava is also given to some other species in the genus '' Psidium'' such as strawberry guava (''
Psidium cattleyanum ''Psidium cattleyanum (World Plants : Psidium cattleianum)'', commonly known as Cattley guava, strawberry guava or cherry guava, is a small tree (2–6 m tall) in the Myrtaceae (myrtle) family. The species is named in honour of English hor ...
'') and to the pineapple guava, ''
Feijoa sellowiana ''Feijoa sellowiana'' is a species of flowering plant in the myrtle family, Myrtaceae. It is native to the highlands of southern Brazil, eastern Paraguay, Uruguay, northern Argentina, and Colombia. It is widely cultivated as an ornamental tree ...
''. In 2019, 55 million tonnes of guavas were produced worldwide, led by India with 45% of the total. Botanically, guavas are berries.


Types

The most frequently eaten species, and the one often simply referred to as "the guava", is the apple guava (''
Psidium guayava ''Psidium guajava'', the common guava, yellow guava, lemon guava, or apple guava is an evergreen shrub or small tree native to the Caribbean, Central America and South America. It is easily pollinated by insects; when cultivated, it is pollina ...
''). Guavas are typical
Myrtoideae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All spe ...
, with tough dark heavy leaves that are opposite, simple, elliptic to ovate, and long. The flowers are white, with five petals and numerous
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s. The fruits are many-seeded berries.


Etymology

The term ''guava'' appears to have been in use since the mid-
16th century The 16th century begins with the Julian year 1501 ( MDI) and ends with either the Julian or the Gregorian year 1600 ( MDC) (depending on the reckoning used; the Gregorian calendar introduced a lapse of 10 days in October 1582). The 16th cent ...
. The name derived from the Taíno, a language of the Awawaks as for ''guava tree'' via the Spanish for . It has been adapted in many European and Asian languages, having a similar form.


Origin and distribution

Guavas originated from an area thought to extend from Mexico, Central America or northern South America throughout the Caribbean region. Archaeological sites in Peru yielded evidence of guava cultivation as early as 2500 BCE. Guava was adopted as a crop in subtropical and tropical Asia, parts of the United States (from Tennessee and North Carolina, southward, as well as the west and Hawaii), tropical Africa, South Asia, Southeast Asia, and Oceania. Guavas were introduced to Florida, US in the 19th century and are grown there as far north as Sarasota, Chipley,
Waldo Waldo may refer to: People * Waldo (given name), a list of people and fictional characters * Waldo (surname), a list of people * Waldo (footballer) (1934-2019), full name Waldo Machado da Silva, Brazilian footballer Places Canada * Waldo, ...
and Fort Pierce. However, they are a primary host of the Caribbean fruit fly and must be protected against infestation in areas of Florida where this pest is present. Guavas are cultivated in many tropical and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
countries. Several species are grown commercially; apple guava and its cultivars are those most commonly traded internationally. Guavas also grow in southwestern Europe, specifically the
Costa del Sol The Costa del Sol (literally "Coast of the Sun" or "Sun Coast") is a region in the south of Spain in the autonomous community of Andalusia, comprising the coastal towns and communities along the coastline of the Province of Málaga and the easte ...
on
Málaga Málaga (, ) is a municipality of Spain, capital of the Province of Málaga, in the autonomous community of Andalusia. With a population of 578,460 in 2020, it is the second-most populous city in Andalusia after Seville and the sixth most pop ...
, (Spain) and Greece where guavas have been commercially grown since the middle of the 20th century and they proliferate as cultivars. Mature trees of most species are fairly cold-hardy and can survive temperatures slightly colder than for short periods of time, but younger plants will likely freeze to the ground. Guavas are of interest to home growers in subtropical areas as one of the few tropical fruits that can grow to fruiting size in pots indoors. When grown from seed, guava trees can bear fruit in two years, and can continue to do so for forty years.


Ecology

''Psidium'' species are eaten by the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
, mainly moths like the Ello Sphinx ('' Erinnyis ello''), ''
Eupseudosoma aberrans ''Eupseudosoma aberrans'' is a moth of the family Erebidae first described by William Schaus in 1905. It is found in Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North Amer ...
'', '' E. involutum'', and '' Hypercompe icasia''. Mites, like ''
Pronematus pruni ''Pronematus pruni'' is a species of mite belonging to the family Tydeidae. This small oval mite is around 300 μm in length with a smooth body and legs much shorter than the body. It has been recorded on ''Prunus domestica'' and ''Psidium guaj ...
'' and ''
Tydeus munsteri ''Tydeus munsteri'' is a species of mite belonging to the family Tydeidae. This very small oval, eyeless mite is around 250 μm in length with a soft body covered in striations. It has been recorded from various plants in the vicinity of Munster ...
'', are known to be crop
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
of the
apple guava ''Psidium guajava'', the common guava, yellow guava, lemon guava, or apple guava is an evergreen shrub or small tree native to the Caribbean, Central America and South America. It is easily pollinated by insects; when cultivated, it is pollina ...
('' P. guajava'') and perhaps other species. The bacterium ''
Erwinia psidii ''Erwinia psidii'' is a Gram-negative bacterium and a phytopathogen of the common guava (''Psidium guajava''), causing rot in branches, flowers and fruits. Recently, it was demonstrated that this species produces two acyl homoserine lactones (' ...
'' causes rot diseases of the apple guava. The fruit is cultivated and favored by humans, and many other animals such as birds consume it, readily dispersing the seeds in their droppings. In Hawaii, strawberry guava ('' P. littorale'') has become an aggressive
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
threatening extinction to more than 100 other plant species. By contrast, several guava species have become rare due to habitat destruction and at least one (Jamaican guava, '' P. dumetorum''), is already extinct. Guava wood is used for meat smoking in Hawaii, and is used at barbecue competitions across the United States. In Cuba and Mexico, the leaves are used in barbecues.


Fruit

Guava fruits, usually long, are round or oval depending on the species. They have a pronounced and typical fragrance, similar to lemon rind but less sharp. The outer skin may be rough, often with a bitter taste, or soft and sweet. Varying between species, the skin can be any thickness, is usually green before maturity, but may be yellow, maroon, or green when ripe. The pulp inside may be sweet or sour and off-white ("white" guavas) to deep pink ("red" guavas). The seeds in the central pulp vary in number and hardness, depending on species.


Production

In 2019, world production of guavas was 55 million tonnes, led by India with 45% of the total (table). Other major producers were China and Thailand.


Culinary uses

In Mexico and other Latin American countries, the popular beverage agua fresca is often made with guava. The entire fruit is a key ingredient in punch, and the juice is often used in culinary sauces (hot or cold), ales, candies, dried snacks, fruit bars, and desserts, or dipped in
chamoy Chamoy may refer to: * Chamoy (sauce), a Mexican condiment * Chamoy, Aube, France * Chamoy Thipyaso (born 1940), Thai prisoner See also * Chamois (disambiguation) {{disambiguation ...
. '' Pulque de guayaba'' ("guayaba" is Spanish for guava) is a popular alcoholic beverage in these regions. In many countries, guava is eaten raw, typically cut into quarters or eaten like an apple; it is also eaten with a pinch of salt and pepper, cayenne powder or a mix of spices (masala). In the Philippines, ripe guava is used in cooking '' sinigang''. Guava is a popular snack in Cuba as ''pastelitos de guayaba''; and in Taiwan, sold on many street corners and night markets during hot weather, accompanied by packets of dried plum powder mixed with sugar and salt for dipping. In east Asia, guava is commonly eaten with sweet and sour dried plum powder mixtures. Guava juice is popular in many countries. The fruit is also often included in fruit salads. Because of its high level of
pectin Pectin ( grc, πηκτικός ': "congealed" and "curdled") is a heteropolysaccharide, a structural acid contained in the primary lamella, in the middle lamella, and in the cell walls of terrestrial plants. The principal, chemical component of ...
, guavas are extensively used to make candies, preserves, jellies,
jams Jams or JAMS may refer to: *Plural form of jam, a type of fruit preserve *Jams (clothing line) *JAMS (organization), United States organization that provides alternative dispute resolution services *The JAMs, former name of The KLF, a British band ...
, and marmalades (such as Brazilian '' goiabada'' and Colombian and Venezuelan '' bocadillo''), and as a marmalade jam served on toast. Red guavas can be used as the base of salted products such as sauces, substituting for tomatoes, especially to minimize the acidity. A drink may be made from an infusion of guava fruits and leaves, which in Brazil is called ''chá-de-goiabeira'', i.e., "tea" of guava tree leaves, considered medicinal.


Constituents


Nutrients

Guavas are rich in
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
and vitamin C, with moderate levels of folic acid (nutrition table). Low in food energy per typical serving, and with few
essential nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
s, a single common guava ('' P. guajava'') fruit contains 257% of the Daily Value (DV) for vitamin C (table). Nutrient content varies across guava cultivars. Although the
strawberry guava ''Psidium cattleyanum (World Plants : Psidium cattleianum)'', commonly known as Cattley guava, strawberry guava or cherry guava, is a small tree (2–6 m tall) in the Myrtaceae (myrtle) family. The species is named in honour of English hor ...
(''P. littorale'' var. ''cattleianum'') has only 39% of the vitamin C in common varieties, its content in a 100 gram serving (90 mg) still provides 100% of the DV.


Phytochemicals

Guava leaves contain both
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
s and polyphenols like (+)-gallocatechin and leucocyanidin. As some of these phytochemicals produce the fruit skin and flesh color, guavas that are red-orange tend to have more polyphenol and carotenoid content than yellow-green ones.


Guava seed oil

Guava seed oil, which may be used for
culinary Culinary arts are the cuisine arts of outline of food preparation, food preparation, cooking and food presentation, presentation of food, usually in the form of meals. People working in this field – especially in establishments such as res ...
or cosmetics products, is a source of beta carotene, vitamin A, vitamin C, copper, zinc and selenium, and is particularly rich in linoleic acid. The composition of fatty acids in guava seed oil is presented in the following table:


Folk medicine

Since the 1950s, guavas – particularly the leaves – have been studied for their constituents, potential biological properties and history in folk medicine.


Parasites

Guavas are one of the most common hosts for fruit flies like '' A. suspensa'', which lay their eggs in overripe or spoiled guavas. The larvae of these flies then consume the fruit until they can proceed into the pupa stage. This
parasitism Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
has led to millions in economic losses for nations in Central America.


See also

*
Myrteae Myrteae is the largest tribe in the plant family Myrtaceae. It includes most of the species of the family that have fleshy fruits. Well-known members include edible fruit such as feijoa (''Feijoa sellowiana''), guava (''Psidium guajava''), jab ...
, the tribe containing guava and closely related plants with fleshy fruit


References

{{Reflist Berries Edible fruits Mexican cuisine Cuban cuisine Belizean cuisine