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The grivet (''Chlorocebus aethiops'') is an
Old World monkey Old World monkey is the common English name for a family of primates known taxonomically as the Cercopithecidae (). Twenty-four genera and 138 species are recognized, making it the largest primate family. Old World monkey genera include baboons ...
with long white tufts of hair along the sides of its face. Some authorities consider this and all of the members of the genus ''
Chlorocebus ''Chlorocebus'' is a genus of medium-sized primates from the family of Old World monkeys. Six species are currently recognized, although some people classify them all as a single species with numerous subspecies. Either way, they make up the ent ...
'' to be a single species, ''Cercopithecus aethiops''. As here defined, the grivet is restricted to
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
,
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
,
Djibouti Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
, and
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopi ...
. In the southern part of its range, it comes into contact with the closely related
vervet monkey The vervet monkey (''Chlorocebus pygerythrus''), or simply vervet, is an Old World monkey of the family Cercopithecidae native to Africa. The term "vervet" is also used to refer to all the members of the genus ''Chlorocebus''. The five distinct ...
(''C. pygerythrus'') and Bale Mountains vervet (''C. djamdjamensis'').
Hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
between them is possible, and may present a threat to the vulnerable Bale Mountains vervet. Unlike that species, the grivet is common and rated as
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
.


Physical description

The grivet's facial skin, hands, and feet are black. The face has a white line above the eyes. It has long, white whiskers on the cheeks. The fur on the back has an olive color, while the front is white. The skin on the stomach has a blue tint. The fur has a bristly feel. The approximate head and body length for males is and for females. The length of the tail for males is about . The body mass ranges from , with females at the smaller end of the scale.


Habitat and distribution

The main habitat of the grivet is savanna woodlands. Its range is
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
east of the
White Nile The White Nile ( ar, النيل الأبيض ') is a river in Africa, one of the two main tributaries of the Nile, the other being the Blue Nile. The name comes from the clay sediment carried in the water that changes the water to a pale color ...
,
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopi ...
, and
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
east to the
Rift Valley A rift valley is a linear shaped lowland between several highlands or mountain ranges created by the action of a geologic rift. Rifts are formed as a result of the pulling apart of the lithosphere due to extensional tectonics. The linear d ...
. It is also found in
Djibouti Djibouti, ar, جيبوتي ', french: link=no, Djibouti, so, Jabuuti officially the Republic of Djibouti, is a country in the Horn of Africa, bordered by Somalia to the south, Ethiopia to the southwest, Eritrea in the north, and the Red ...
. The grivet needs to live around a source of water, especially during the dry season. It is able to adapt to many environments.


Local and indigenous names

In
Tigrinya language (; also spelled Tigrigna) is an Ethio-Semitic language commonly spoken Eritrea and in northern Ethiopia's Tigray Region by the Tigrinya and Tigrayan peoples. It is also spoken by the global diaspora of these regions. History and literat ...
: (wi’ag)


Behavior

The grivet is most active in the morning and early evening. It stays on the ground most of the day to eat, and at night it sleeps in trees. The grivet spends a lot of time grooming, playing, climbing, and play fighting; all of these things help to ensure its survival. Its eating habits consist of eating mostly fruits, vegetables, and sometimes small mammals, insects, and birds, making it an
omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nu ...
. It also scavenges for human food. It must drink water daily, especially in the dry seasons. It is one of few species that has multiple-male groups that are of moderate size. In the hierarchy of males, an individual shows his dominance by putting his tail in a stiff, upright position and strolling past lower-ranked males. They travel in packs, and usually move on all fours or
quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...
ally, except when using both hands for carrying, when they manage to walk and run quite comfortably on two legs. Groups can range from five to over 70. Females will have a limited number of mates, while males may have several. Swelling of the female's vulva alerts males as to when she is in
estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...
. Giving birth to one baby at a time is common, and gestation usually lasts 2-3 months. When the baby is born, the mother cleans the infant and bites off the umbilical cord. Young have pink faces and black hair. Around two months are needed for them to get their adult coats. The first few months, the infant stays very close to its mother, but after 6 months, the infant is weaned.


Conservation

Grivets are occasionally hunted as
bushmeat Bushmeat is meat from wildlife species that are hunted for human consumption, most often referring to the meat of game in Africa. Bushmeat represents a primary source of animal protein and a cash-earning commodity for inhabitants of humid tropi ...
. They are killed for either commercial or subsistence purposes. Although not endangered, they are threatened through destruction of habitat - forests. They are preyed on by large snakes, leopards, humans, and sometimes baboons. Grivets may live for 13 years.


Relationship with humans

The grivet is one of five species of monkeys known to have been kept in ancient Egypt, the others being the
hamadryas baboon The hamadryas baboon (''Papio hamadryas'' ) is a species of baboon within the Old World monkey family. It is the northernmost of all the baboons, being native to the Horn of Africa and the southwestern region of the Arabian Peninsula. These re ...
, the
olive baboon The olive baboon (''Papio anubis''), also called the Anubis baboon, is a member of the family Cercopithecidae Old World monkeys. The species is the most wide-ranging of all baboons, being native to 25 countries throughout Africa, extending fr ...
, the
patas monkey The common patas monkey (''Erythrocebus patas''), also known as the wadi monkey or hussar monkey, is a ground-dwelling monkey distributed over semi-arid areas of West Africa, and into East Africa. Taxonomy There is some confusion surrounding ...
, and the
barbary macaque The Barbary macaque (''Macaca sylvanus''), also known as Barbary ape, is a macaque species native to the Atlas Mountains of Algeria, Libya, Tunisia and Morocco, along with a small introduced population in Gibraltar. It is the type species of the ...
. Grivets were imported from the
Land of Punt The Land of Punt ( Egyptian: '' pwnt''; alternate Egyptological readings ''Pwene''(''t'') /pu:nt/) was an ancient kingdom known from Ancient Egyptian trade records. It produced and exported gold, aromatic resins, blackwood, ebony, ivory an ...
, as attested in paintings and in the '' Tale of the Shipwrecked Sailor''. They were sometimes traded as far afield as
Assyria Assyria ( Neo-Assyrian cuneiform: , romanized: ''māt Aššur''; syc, ܐܬܘܪ, ʾāthor) was a major ancient Mesopotamian civilization which existed as a city-state at times controlling regional territories in the indigenous lands of the A ...
. They are rarer in representations than baboons, and unlike baboons, do not seem to have borne individual names.Dieter Kessler, "Monkeys and Baboons", in Donald B. Redford (ed.), ''The Oxford Encyclopedia of Ancient Egypt'' (Oxford University Press, 2001 nline 2005, retrieved 11 March 2019. Grivets are depicted on Egyptian tombs as house pets and on leashes. In some depictions, they may symbolize male sexuality. Early Dynastic statuettes of grivets have been found in sanctuaries, where they may have been votive offerings to the baboon god. A grivet shooting a bow was an aspect of the invisible god
Atum Atum (, Egyptian: ''jtm(w)'' or ''tm(w)'', ''reconstructed'' ; Coptic ''Atoum''), sometimes rendered as Atem or Tem, is an important deity in Egyptian mythology. Name Atum's name is thought to be derived from the verb ''tm'' which means 'to co ...
, and at Deltaic Babylon, a grivet was the town god represented by a statue in the temple. Since the 1960s, the grivet has been harvested on an industrial scale by humans for their kidneys, which are used in poliovirus and adenovirus vaccine production. In 1967, the Marburg virus in grivets caused the death of 31 vaccine manufacturers. In the medical literature, the grivet and the green monkey are often confused. They are also used in medical research such as the testing of dengue vaccines as well as in the testing of human cosmetics.


References

{{Taxonbar, from=Q257470 Chlorocebus Fauna of East Africa Mammals of Djibouti Mammals of South Sudan Mammals of Sudan Mammals of Eritrea Mammals of Ethiopia Mammals described in 1758 Taxa named by Carl Linnaeus Land of Punt