HOME

TheInfoList



OR:

The greylag goose or graylag goose (''Anser anser'') is a species of large
goose A goose ( : geese) is a bird of any of several waterfowl species in the family Anatidae. This group comprises the genera ''Anser'' (the grey geese and white geese) and '' Branta'' (the black geese). Some other birds, mostly related to the she ...
in the waterfowl family
Anatidae The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating on ...
and the type species of the genus ''
Anser Anser may refer to: People * Anser (poet), poet of ancient Rome * Anser Farooq, Canadian lawyer Other uses * ANSER, a security and defense analysis group * ''Anser'' (bird), a genus of geese * Anser (putter), a model of golf club made by ...
''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A large bird, it measures between in length, with an average weight of . Its distribution is widespread, with birds from the north of its range in Europe and Asia migrating southwards to spend the winter in warmer places. It is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
''
Anser Anser may refer to: People * Anser (poet), poet of ancient Rome * Anser Farooq, Canadian lawyer Other uses * ANSER, a security and defense analysis group * ''Anser'' (bird), a genus of geese * Anser (putter), a model of golf club made by ...
'' and is the ancestor of most breeds of
domestic goose A domestic goose is a goose that humans have domesticated and kept for their meat, eggs, or down feathers. Domestic geese have been derived through selective breeding from the wild greylag goose (''Anser anser domesticus'') and swan goose (''Ans ...
, having been domesticated at least as early as 1360 BC. The genus name is from ''anser'', the Latin for "goose". Greylag geese travel to their northerly breeding grounds in spring, nesting on moorlands, in marshes, around lakes and on coastal islands. They normally mate for life and nest on the ground among vegetation. A clutch of three to five eggs is laid; the female incubates the eggs and both parents defend and rear the young. The birds stay together as a family group, migrating southwards in autumn as part of a flock, and separating the following year. During the winter they occupy semi-aquatic habitats, estuaries, marshes and flooded fields, feeding on grass and often consuming agricultural crops. Some populations, such as those in southern England and in urban areas across the species' range, are primarily resident and occupy the same area year-round.


Taxonomy

''Anser anser'', the greylag goose, is a member of the waterfowl family
Anatidae The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating on ...
. It was first described by the Swedish naturalist Carl Linnaeus in 1758 as ''Anas anser'', but was transferred two years later to the new genus ''
Anser Anser may refer to: People * Anser (poet), poet of ancient Rome * Anser Farooq, Canadian lawyer Other uses * ANSER, a security and defense analysis group * ''Anser'' (bird), a genus of geese * Anser (putter), a model of golf club made by ...
'', erected by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
, where it is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised; ''A. a. anser'', the western greylag goose, which breeds in Iceland and northern and central Europe; ''A. a. rubrirostris'', the eastern greylag goose, which breeds in Romania, Turkey, and Russia eastwards to northeastern China. The two subspecies
intergrade In zoology, intergradation is the way in which two distinct subspecies are connected via areas where populations are found that have the characteristics of both. There are two types of intergradation: primary and secondary intergradation. Primary ...
where their ranges meet. The greylag goose sometimes hybridises with other species of goose, including the
barnacle goose The barnacle goose (''Branta leucopsis'') is a species of goose that belongs to the genus ''Branta'' of black geese, which contains species with largely black plumage, distinguishing them from the grey '' Anser'' species. Despite its superficial ...
(''Branta leucopsis'') and the
Canada goose The Canada goose (''Branta canadensis''), or Canadian goose, is a large wild goose with a black head and neck, white cheeks, white under its chin, and a brown body. It is native to the arctic and temperate regions of North America, and it is ...
(''Branta canadensis''), and occasionally with the
mute swan The mute swan (''Cygnus olor'') is a species of swan and a member of the waterfowl family Anatidae. It is native to much of Eurosiberia, and (as a rare winter visitor) the far north of Africa. It is an introduced species in North America, home ...
(''Cygnus olor''). The greylag goose was one of the first animals to be domesticated; this happened at least 3,000 years ago in Ancient Egypt, the domestic subspecies being known as ''A. a. domesticus''. As the domestic goose is a subspecies of the greylag goose they are able to interbreed, with the offspring sharing characteristics of both wild and domestic birds.


Description

The greylag is the largest and bulkiest of the grey geese of the genus ''Anser'', but is more lightly built and agile than its domestic relative. It has a rotund, bulky body, a thick and long neck, and a large head and bill. It has pink legs and feet, and an orange or pink bill with a white or brown
nail Nail or Nails may refer to: In biology * Nail (anatomy), toughened protective protein-keratin (known as alpha-keratin, also found in hair) at the end of an animal digit, such as fingernail * Nail (beak), a plate of hard horny tissue at the tip ...
(hard horny material at tip of upper mandible). It is long with a wing length of . It has a tail , a bill of long, and a tarsus of . It weighs , with a mean weight of around . The wingspan is . Males are generally larger than females, with the
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most anim ...
more pronounced in the eastern subspecies ''rubirostris'', which is larger than the nominate subspecies on average. The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of the greylag goose is greyish brown, with a darker head and paler breast and belly with a variable amount of black spotting. It has a pale grey forewing and
rump Rump may refer to: * Rump (animal) ** Buttocks * Rump steak, slightly different cuts of meat in Britain and America * Rump kernel, software run in userspace that offers kernel functionality in NetBSD Politics * Rump cabinet *Rump legislature * R ...
which are noticeable when the bird is in flight or stretches its wings on the ground. It has a white line bordering its upper flanks, and its
wing coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
are light coloured, contrasting with its darker flight feathers. Its plumage is patterned by the pale fringes of the feathers. Juveniles differ mostly in their lack of black speckling on the breast and belly and by their greyish legs. Adults have a distinctive 'concertina' pattern of folds in the feathers on their necks. The greylag goose has a loud cackling call similar to that of the domestic goose, "aahng-ung-ung", uttered on the ground or in flight. There are various subtle variations used under different circumstances, and individual geese seem to be able to identify other known geese by their voices. The sound made by a flock of geese resembles the baying of hounds. Goslings chirp or whistle lightly, and adults hiss if threatened or angered.


Distribution and habitat

This species has a
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Si ...
distribution. The nominate subspecies breeds in Iceland, Norway, Sweden, Finland, the Baltic States, northern Russia, Poland, eastern Hungary and Romania. It also breeds locally in the United Kingdom, Denmark, Germany, Austria, the Czech Republic, Slovakia and North Macedonia. The eastern race extends eastwards across a broad swathe of Asia to China. European birds migrate southwards to the Mediterranean region and North Africa. Asian birds migrate to Azerbaijan, Iran, Pakistan, northern India, Bangladesh and eastward to China. In North America, there are both
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
domestic geese, which are similar to greylags, and occasional
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, tempora ...
greylags. Greylag geese seen in the wild in New Zealand probably originated from the escape of farmyard geese, and a similar situation has occurred in Australia, where feral birds are now established in the east and southeast of the country. In their breeding quarters, they are found on moors with scattered lochs, in marshes, fens and peat-bogs, besides lakes and on little islands some way out to sea. They like dense ground cover of reeds, rushes, heather, bushes and willow thickets. In their winter quarters, they frequent salt marshes, estuaries, freshwater marshes, steppes, flooded fields, bogs and pasture near lakes, rivers and streams. They also visit agricultural land where they feed on winter cereals, rice, beans or other crops, moving at night to shoals and sand-banks on the coast, mud-banks in estuaries or secluded lakes. Large numbers of immature birds congregate each year to moult on the Rone Islands near Gotland in the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...
. Since the 1950s, increases in winter temperatures have resulted in greylag geese, breeding in central Europe, reducing their winter migration distances. Wintering grounds closer to home can therefore be exploited, meaning that the geese can return to set up breeding territories earlier the following spring. In Great Britain, their numbers had declined as a breeding bird, retreating north to breed wild only in the
Outer Hebrides The Outer Hebrides () or Western Isles ( gd, Na h-Eileanan Siar or or ("islands of the strangers"); sco, Waster Isles), sometimes known as the Long Isle/Long Island ( gd, An t-Eilean Fada, links=no), is an island chain off the west coas ...
and the northern mainland of Scotland. However, during the 20th century, feral populations have been established elsewhere, and they have now re-colonised much of England. These populations are increasingly coming into contact and merging. The greylag goose has become a
pest species A pest is any animal or plant harmful to humans or human concerns. The term is particularly used for creatures that damage crops, livestock, and forestry or cause a nuisance to people, especially in their homes. Humans have modified the environ ...
in several areas where its population has increased sharply. In Norway, the number of greylag geese is estimated to have increased three- to fivefold between 1995 and 2015. As a consequence, farmers' problems caused by goose grazing on farmland have increased considerably. This problem is also evident for the
pink-footed goose The pink-footed goose (''Anser brachyrhynchus'') is a goose which breeds in eastern Greenland, Iceland and Svalbard. It is migratory, wintering in northwest Europe, especially Ireland, Great Britain, the Netherlands, and western Denmark. The nam ...
. In the
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) nort ...
islands the population has increased dramatically: there were 300 breeding pairs, increasing to 10,000 in 2009, and 64,000 in 2019. Due to extensive damage caused to crops, the hunting season for the greylag goose in the Orkney islands is now most of the year.


Behaviour

Greylag geese are largely
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
and feed chiefly on
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
es. Short, actively growing grass is more nutritious and greylag geese are often found grazing in pastures with sheep or cows. Because of its low nutrient status, they need to feed for much of their time; the herbage passes rapidly through the gut and is voided frequently. The tubers of sea clubrush (''Bolboschoenus maritimus'') are also taken as well as berries and water plants such as
duckweed Lemnoideae is a subfamily of flowering aquatic plants, known as duckweeds, water lentils, or water lenses. They float on or just beneath the surface of still or slow-moving bodies of fresh water and wetlands. Also known as bayroot, they arose fr ...
(''Lemna'') and floating sweetgrass (''Glyceria fluitans''). In wintertime they eat grass and leaves but also glean grain on cereal stubbles and sometimes feed on growing crops, especially during the night. They have been known to feed on
oat The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural, unlike other cereals and pseudocereals). While oats are suitable for human con ...
s,
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeological ...
, barley,
buckwheat Buckwheat (''Fagopyrum esculentum''), or common buckwheat, is a flowering plant in the knotweed family Polygonaceae cultivated for its grain-like seeds and as a cover crop. The name "buckwheat" is used for several other species, such as ''Fagop ...
,
lentil The lentil (''Lens culinaris'' or ''Lens esculenta'') is an edible legume. It is an annual plant known for its lens-shaped seeds. It is about tall, and the seeds grow in pods, usually with two seeds in each. As a food crop, the largest produ ...
s,
pea The pea is most commonly the small spherical seed or the seed-pod of the flowering plant species ''Pisum sativum''. Each pod contains several peas, which can be green or yellow. Botanically, pea pods are fruit, since they contain seeds and d ...
s and
root crop Root vegetables are underground plant parts eaten by humans as food. Although botany distinguishes true roots (such as taproots and tuberous roots) from non-roots (such as bulbs, corms, rhizomes, and tubers, although some contain both hypocotyl a ...
s.
Acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s are sometimes consumed, and on the coast,
seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families ( Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the ...
(''Zostera'' sp.) may be eaten. In the 1920s in Britain, the pink-footed goose "discovered" that potatoes were edible and started feeding on waste potatoes. The greylag followed suit in the 1940s and now regularly searches for tubers on ploughed fields. They also consume small
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
,
amphibians Amphibians are four-limbed and ectothermic vertebrates of the class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial, fossorial, arbore ...
,
crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group c ...
,
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is estim ...
and
insects Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of ...
. Greylag geese tend to pair bond in long-term
monogamous Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time (serial monogamy) — as compared to the various forms of non-monogamy (e.g., polyg ...
relationships. Most such pairs are probably life-long partnerships, though 5 to 8% of the pairs separate and re-mate with other geese. Birds in heterosexual pairs may engage in promiscuous behavior, despite the opposition of their mates. Homosexual pairs are common (14 to 20% of the pairs may be ganders, depending on flock), and share the characteristics of heterosexual pairs with the exceptions that the bonds appear to be closer, based on the intensity of their displays. Same-sex pairs also engage in courtship and sexual relations, and often assume high-ranking positions in the flock as a result of their superior strength and courage, leading some to speculate that they may serve as guardians of the flock. The sexual preference of the birds is generally flexible, as more than half of widowers re-pair with a bird of the opposite sex. The nest is on the ground among heather, rushes, dwarf shrubs or reeds, or on a raft of floating vegetation. It is built from pieces of reed, sprigs of heather, grasses and moss, mixed with small feathers and down. A typical clutch is four to six eggs, but fewer eggs or larger numbers are not unusual. The eggs are creamy-white at first but soon become stained, and average . They are mostly laid on successive days and incubation starts after the last one is laid. The female does the incubation, which lasts about twenty-eight days, while the male remains on guard somewhere near. The chicks are
precocial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
and able to leave the nest soon after hatching. Both parents are involved in their care and they soon learn to peck at food and become fully-fledged at eight or nine weeks, about the same time as their parents regain their ability to fly after moulting their main wing and tail feathers a month earlier. Immature birds undergo a similar moult, and move to traditional, safe locations before doing so because of their vulnerability while flightless. Greylag geese are
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
birds and form flocks. This has the advantage for the birds that the vigilance of some individuals in the group allows the rest to feed without having to constantly be alert to the approach of
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
. After the eggs hatch, some grouping of families occur, enabling the geese to defend their young by their joint actions, such as mobbing or attacking predators. After driving off a predator, a gander will return to its mate and give a "triumph call", a resonant honk followed by a low-pitched cackle, uttered with neck extended forward parallel with the ground. The mate and even unfledged young reciprocate in kind. Young greylags stay with their parents as a family group, migrating with them in a larger flock, and only dispersing when the adults drive them away from their newly established breeding
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
the following year. At least in Europe, patterns of migration are well understood and follow traditional routes with known staging sites and wintering sites. The young learn these locations from their parents which normally stay together for life. Greylags leave their northern breeding areas relatively late in the autumn, for example completing their departure from Iceland by November, and start their return migration as early as January. Birds that breed in Iceland overwinter in the British Isles; those from Central Europe overwinter as far south as Spain and North Africa; others migrate down to the Balkans, Turkey and Iraq for the winter. stated to be from


In human culture

The greylag was once revered across Eurasia. It was linked with the goddess of healing, Gula, a forerunner of the
Sumeria Sumer () is the earliest known civilization in the historical region of southern Mesopotamia (south-central Iraq), emerging during the Chalcolithic and early Bronze Ages between the sixth and fifth millennium BC. It is one of the cradles of ...
n fertility goddess
Ishtar Inanna, also sux, 𒀭𒊩𒌆𒀭𒈾, nin-an-na, label=none is an List of Mesopotamian deities, ancient Mesopotamian goddess of love, war, and fertility. She is also associated with beauty, sex, Divine law, divine justice, and political p ...
, in the cities of the
Tigris The Tigris () is the easternmost of the two great rivers that define Mesopotamia, the other being the Euphrates. The river flows south from the mountains of the Armenian Highlands through the Syrian and Arabian Deserts, and empties into the ...
-
Euphrates The Euphrates () is the longest and one of the most historically important rivers of Western Asia. Together with the Tigris, it is one of the two defining rivers of Mesopotamia ( ''the land between the rivers''). Originating in Turkey, the Euph ...
delta over 5,000 years ago. In Ancient Egypt, geese symbolised the sun god Ra. In
Ancient Greece Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cult ...
and
Rome , established_title = Founded , established_date = 753 BC , founder = King Romulus (Romulus and Remus, legendary) , image_map = Map of comune of Rome (metropolitan city of Capital Rome, region Lazio, Italy).svg ...
, they were associated with the goddess of love,
Aphrodite Aphrodite ( ; grc-gre, Ἀφροδίτη, Aphrodítē; , , ) is an ancient Greek goddess associated with love, lust, beauty, pleasure, passion, and procreation. She was syncretized with the Roman goddess . Aphrodite's major symbols include ...
, and goose fat was used as an
aphrodisiac An aphrodisiac is a substance that increases sexual desire, sexual attraction, sexual pleasure, or sexual behavior. Substances range from a variety of plants, spices, foods, and synthetic chemicals. Natural aphrodisiacs like cannabis or cocain ...
. Since they were sacred birds, they were kept on Rome's Capitoline Hill, from where they raised the alarm when the Gauls attacked in 390 BC. The goose's role in fertility survives in modern British tradition in the
nursery rhyme A nursery rhyme is a traditional poem or song for children in Britain and many other countries, but usage of the term dates only from the late 18th/early 19th century. The term Mother Goose rhymes is interchangeable with nursery rhymes. From t ...
'' Goosey Goosey Gander'', which preserves its sexual overtones ("And in my lady's chamber"), while "to goose" still has a sexual meaning. The tradition of pulling a wishbone derives from the tradition of eating a roast goose at
Michaelmas Michaelmas ( ; also known as the Feast of Saints Michael, Gabriel, and Raphael, the Feast of the Archangels, or the Feast of Saint Michael and All Angels) is a Christian festival observed in some Western liturgical calendars on 29 September, a ...
, where the goose bone was once believed to have the powers of an oracle. For that festival, in
Thomas Bewick Thomas Bewick (c. 11 August 17538 November 1828) was an English wood-engraver and natural history author. Early in his career he took on all kinds of work such as engraving cutlery, making the wood blocks for advertisements, and illustrating ch ...
's time, geese were driven in thousand-strong flocks on foot from farms all over the East of England to London's
Cheapside Cheapside is a street in the City of London, the historic and modern financial centre of London, which forms part of the A40 London to Fishguard road. It links St. Martin's Le Grand with Poultry. Near its eastern end at Bank junction, where ...
market, covering some per day. Some farmers painted the geese's feet with tar and sand to protect them from road wear as they walked. Greylag geese were domesticated by at least 1360 BC, when images of domesticated birds resembling the eastern race, ''Anser anser rubirostris'' (which like modern farmyard geese, but unlike western greylags, have a pink beak) were painted in Ancient Egypt. Goose feathers were used as quill pens, the best being the primary feathers of the left-wing, whose "curvature bent away from the eyes of right-handed writers". The feathers also served to fletch arrows. In
ethology Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and object ...
, the greylag goose was the subject of
Konrad Lorenz Konrad Zacharias Lorenz (; 7 November 1903 – 27 February 1989) was an Austrian zoologist, ethologist, and ornithologist. He shared the 1973 Nobel Prize in Physiology or Medicine with Nikolaas Tinbergen and Karl von Frisch. He is often regarded ...
's pioneering studies of imprinting
behaviour Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
.


Gallery

Greylag goose (anser anser).JPG, Variant with white "front" Anser anser EM1B5927 (36071281286).jpg, In flight Birds of Sweden 2016 44.jpg, Swimming Ánsar común (Anser anser) en el Palacio de Nymphenburg, Múnich, Alemania08.JPG, Detail Greylag Goose from the Crossley ID Guide Britain and Ireland.jpg, ID composite Grågås (Anser-anser)-Ystad-2015.jpg, Flock taking off over water Anser anser MWNH 0955.JPG, Egg, Collection
Museum Wiesbaden The Museum Wiesbaden is a two-branch museum of art and natural history in the Hessian capital of Wiesbaden, Germany. It is one of the three Hessian State museums, in addition to the museums in Kassel and Darmstadt. History The foundation o ...
Anser anser EM1B9595 (33952161753).jpg, Female with chicks Alerte grauwe ganzen-4961601.webm, On
Texel Texel (; Texels dialect: ) is a municipality and an island with a population of 13,643 in North Holland, Netherlands. It is the largest and most populated island of the West Frisian Islands in the Wadden Sea. The island is situated north of Den ...
, Netherlands Grågås (Anser anser) Ystad 2016.ogv, In
Ystad Ystad (; older da, Ysted) is a town and the seat of Ystad Municipality, in Scania County, Sweden. Ystad had 18,350 inhabitants in 2010. The settlement dates from the 11th century and has become a busy ferryport, local administrative centre, and ...
Greylag-goose-63088.jpg, Full body


References


Further reading

* *


External links


Ageing and sexing (PDF) by Javier Blasco-Zumeta & Gerd-Michael HeinzeGreylag Goose at RSPB A to Z of UK Birds
*
Greylag Goose pictures
Wildlife Greylag Goose photos- adult with nestlings and voice at nature-pictures.org * *
Greylag Goose Educational video from Avi Birds
{{Authority control
greylag goose The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus '' Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A ...
greylag goose The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus '' Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A ...
Birds of Eurasia
greylag goose The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus '' Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A ...
Articles containing video clips
greylag goose The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus '' Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A ...