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botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, a fruit is the
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
-bearing structure in
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s. Edible fruits in particular have long propagated using the movements of humans and animals in a symbiotic relationship that is the means for seed dispersal for the one group and nutrition for the other; in fact, humans and many animals have become dependent on fruits as a source of food. Consequently, fruits account for a substantial fraction of the world's agricultural output, and some (such as the
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ancest ...
and the pomegranate) have acquired extensive cultural and symbolic meanings. In common language usage, "fruit" normally means the seed-associated fleshy structures (or produce) of plants that typically are sweet or sour and edible in the raw state, such as
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ancest ...
s, bananas,
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
s, lemons, oranges, and
strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...
. In botanical usage, the term "fruit" also includes many structures that are not commonly called "fruits" in everyday language, such as nuts, bean pods, corn kernels, tomatoes, and wheat grains.


Botanical vs. culinary

Many common language terms used for fruit and seeds differ from botanical classifications. For example, in botany, a ''fruit'' is a ripened ovary or carpel that contains seeds; e.g., an apple, pomegranate, tomato or a pumpkin. A '' nut'' is a type of fruit (and not a seed), and a ''seed'' is a ripened ovule. In culinary language, a ''fruit'' is the sweet- or not sweet- (even sour-) tasting produce of a specific plant (e.g., a peach, pear or lemon); ''nuts'' are hard, oily, non-sweet plant produce in shells ( hazelnut, acorn). '' Vegetables'', so called, typically are
savory Savory or Savoury may refer to: Common usage * Herbs of the genus ''Satureja'', particularly: ** Summer savory (''Satureja hortensis''), an annual herb, used to flavor food ** Winter savory (''Satureja montana''), a perennial herb, also used to ...
or non-sweet produce ( zucchini, lettuce, broccoli, and tomato); but some may be sweet-tasting (sweet potato). Examples of botanically classified fruit that typically are called vegetables include: cucumber, pumpkin, and squash ( all are cucurbits); beans,
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics, important to both small an ...
s, and peas ( all legumes); corn, eggplant, bell pepper (or sweet pepper), and tomato (see image). The spices chili pepper and allspice are fruits, botanically speaking. In contrast, rhubarb is often called a fruit when used in making pies, but the edible produce of rhubarb is actually the leaf stalk or petiole of the plant. Edible gymnosperm seeds are often given fruit names, e.g., ginkgo nuts and pine nuts. Botanically, a cereal grain, such as corn, rice, or wheat is a kind of fruit (termed a caryopsis). However, the fruit wall is thin and fused to the seed coat, so almost all the edible grain-fruit is actually a seed.


Structure

The outer layer, often edible, of most fruits is called the ''pericarp''. Typically formed from the ovary, it surrounds the seeds; in some species, however, other structural tissues contribute to or form the edible portion. The pericarp may be described in three layers from outer to inner, i.e., the ''epicarp'', ''mesocarp'' and ''endocarp''. Fruit that bears a prominent pointed terminal projection is said to be ''beaked''.


Development

A fruit results from the fertilizing and maturing of one or more flowers. The
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pist ...
, which contains the '' stigma-style-ovary'' system, is centered in the flower-head, and it forms all or part of the fruit —(see graphic: 'the parts of a flower'). Inside the ovary(ies) are one or more ovules. Here begins a complex sequence called '' double fertilization'': a female gametophyte produces an egg cell for the purpose of fertilization. (A female gametophyte is called a
megagametophyte A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the sex ...
, and also called the
embryo sac In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fem ...
.) After double fertilization, the ovules will become seeds. Ovules are fertilized in a process that starts with
pollination Pollination is the transfer of pollen from an Stamen, anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by Anemophily, wind. Pollinating agents can ...
, which is the movement of pollen from the stamens to the stigma-style-ovary system within the flower-head, (see graphic). After pollination, a pollen tube grows from the (deposited) pollen through the stigma down the style into the ovary to the ovule. Two sperm are transferred from the pollen to a megagametophyte. Within the megagametophyte one sperm unites with the egg, forming a
zygote A zygote (, ) is a eukaryotic cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individual organism. In multicell ...
, while the second sperm enters the central cell forming the endosperm mother cell, which completes the double fertilization process. Later the zygote will give rise to the embryo of the seed, and the endosperm mother cell will give rise to endosperm, a nutritive tissue used by the embryo. As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, the ''pericarp'', may become fleshy (as in berries or drupes), or it may form a hard outer covering (as in nuts). In some multiseeded fruits, the extent to which a fleshy structure develops is proportional to the number of fertilized ovules. The pericarp typically is differentiated into two or three distinct layers; these are called the ''exocarp'' (outer layer, also called epicarp), ''mesocarp'' (middle layer), and ''endocarp'' (inner layer) —(see image of apple-section). In some fruits the
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...
s, petals, stamens and/or the style of the flower fall away as the fleshy fruit ripens. However, for simple fruits derived from an ''
inferior ovary In the flowering plants, an ovary is a part of the female reproductive organ of the flower or gynoecium. Specifically, it is the part of the pistil which holds the ovule(s) and is located above or below or at the point of connection with the ...
'' —i.e., one that lies ''below'' the attachment of other floral parts, (see graphic re 'insertion point')— there are parts (including petals, sepals, and stamens) that fuse with the ovary and ripen with it. For such a case, when floral parts other than the ovary form a significant part of the fruit that develops, it is called an accessory fruit. Examples of accessory fruits include apple, rose hip, strawberry, pineapple; see below, and "Table of fleshy fruit examples". Because several parts of the flower besides the ovary may contribute to the structure of a fruit, it is important to study flower structure to understand how a particular fruit forms. There are three general modes of fruit development: * Apocarpous fruits develop from a ''single flower'' (while having one or more separate, unfused, carpels); they are the simple fruits. *
Syncarpous Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils' ...
fruits develop from a ''single
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pist ...
'' (having two or more carpels fused together). * Multiple fruits form from many flowers —i.e., an inflorescence of flowers. File:Nectarine Fruit Development.jpg, The development sequence of a typical drupe, the
nectarine The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, n ...
(''Prunus persica'') over a 7.5 month period, from bud formation in early winter to fruit ripening in midsummer (see image page for further information) File:Mature flower diagram.svg, The parts of a flower, showing the stigma-style-ovary system. File:Pome apples text.jpg, An apple is a simple fleshy fruit. Key parts are the epicarp, or exocarp, or outer skin, (not labelled); and the mezocarp and endocarp (labelled). image:Ovary position.svg, upright 1.5, Insertion point: There are three positions of insertion of the ovary at the base of a flower: I superior; II half-inferior; III inferior. The 'insertion point' is where the
androecium The stamen ( plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fil ...
parts ''(a)'', the petals ''(p)'', and the sepals ''(s)'' all converge and attach to the receptacle ''(r)''. (Ovary= gynoecium ''(g)''.) File:Noni fruit dev.jpg, upright 1.5, In the noni, flowers are produced in time-sequence along the stem. It is possible to see a progression of flowering, fruit development, and fruit ripening.


Classification of fruits

Consistent with the three modes of fruit development plant scientists have classified fruits into three main groups: simple fruits, aggregate fruits, and multiple (or composite) fruits. The groupings reflect how the ovary and other flower organs are arranged and how the fruits develop, but they are not evolutionarily relevant as diverse plant taxa may be in the same group. While the section of a
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately fr ...
that produces spores is called a ''fruiting'' body, fungi are members of the fungi kingdom and not of the plant kingdom.


Simple fruits

Simple fruits are the result of the ripening-to-fruit of a simple or compound ovary in a ''single flower'' with a ''single pistil''. In contrast, a single flower with numerous pistils typically produces an
aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
; and the merging of several flowers, or a 'multiple' of flowers, results in a 'multiple' fruit. A simple fruit is further classified as to whether it is dry or fleshy. To distribute their seeds, dry fruits may split open and discharge their seeds to the winds, which is called dehiscence. Or the distribution process may rely upon the decay and degradation of the fruit to expose the seeds; or it may rely upon the eating of fruit and excreting of seeds by frugivores —both are called indehiscence. Fleshy fruits do not split open, but they also are indehiscent and they may also rely on frugivores for distribution of their seeds. Typically, the entire outer layer of the ovary wall ripens into a potentially edible pericarp. Types of dry simple fruits, (with examples) include: * achene – most commonly seen in aggregate fruits (e.g., strawberry, see below). * capsule – (
Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest ...
: botanically, it is not a nut). * caryopsis – (cereal grains, including wheat, rice, oats, barley). * cypsela – an achene-like fruit derived from the individual florets in a capitulum: ( dandelion). * fibrous drupe – ( coconut, walnut: botanically, neither is a true nut.). * follicle – follicles are formed from a single carpel, and opens by one suture: (
milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to hum ...
); also commonly seen in aggregate fruits: (
magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendr ...
, peony). * legume – ( bean, pea,
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics, important to both small an ...
: botanically, the peanut is the seed of a legume, not a nut). * loment – a type of indehiscent legume: ( sweet vetch or wild potato). * nut – ( beechnut, hazelnut, acorn (of the oak): botanically, these are true nuts). * samara – (
ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non-gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...
, elm, maple key). * schizocarp, see below – ( carrot seed). * silique – ( radish seed). * silicle – ( shepherd's purse). * utricle – ( beet, '' Rumex''). Fruits in which part or all of the ''pericarp'' (fruit wall) is fleshy at maturity are termed ''fleshy simple fruits''. Types of fleshy simple fruits, (with examples) include: * berry – The berry is the most common type of fleshy fruit. The entire outer layer of the ovary wall ripens into a potentially edible "pericarp", (see below). * stone fruit or drupe – The definitive characteristic of a drupe is the hard, "lignified" stone (sometimes called the "pit"). It is derived from the ovary wall of the flower: apricot, cherry, olive,
peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, n ...
, plum, mango. * pome – The pome fruits:
apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple trees are cultivated worldwide and are the most widely grown species in the genus '' Malus''. The tree originated in Central Asia, where its wild ancest ...
s, pears,
rosehip The rose hip or rosehip, also called rose haw and rose hep, is the accessory fruit of the various species of rose plant. It is typically red to orange, but ranges from dark purple to black in some species. Rose hips begin to form after pollin ...
s,
saskatoon berry ''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to North America. Description It is a deciduous shrub or sma ...
, et al., are a
syncarpous Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils' ...
(fused) fleshy fruit, a simple fruit, developing from a half-inferior ovary, see graphic re 'Insertion point'. Pomes are of the family Rosaceae,


Berries

Berries are a type of simple fleshy fruit that issue from a single ovary. (The ovary itself may be compound, with several carpels.) The botanical term "true berry" includes grapes, currants, cucumbers, eggplants (aubergines), tomatoes, chili peppers, and bananas, but excludes certain fruits that are called "-berry" by culinary custom or by common usage of the term —such as strawberries and raspberries. Berries may be formed from one or more carpels (i.e., from the simple or compound ovary) from the same, single flower. Seeds typically are embedded in the fleshy interior of the ovary. Examples here and in the table below: * tomato – In culinary terms, the tomato is regarded as a vegetable, but it is botanically classified as a fruit and a berry. * banana – The fruit has been described as a "leathery berry". In cultivated varieties, the seeds are diminished nearly to non-existence. *
pepo PEPO Lappeenranta (abbreviated PEPO) is a football club from Lappeenranta in Finland. The club was formed in 1958 and their home ground is at the Kimpinen Sports Centre. The men's first team currently plays in the Ykkönen (Second Division). ...
– Berries with skin that is hardened:
cucurbits The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most important to humans are: *''Cucurbita'' – squash, pumpkin, zucchini, some gourds *'' Lagena ...
, including gourds, squash, melons. * hesperidium – Berries with a rind and a juicy interior: most
citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...
fruit. * cranberry, gooseberry,
redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus '' Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribe ...
,
grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
. The ''strawberry'', regardless of its appearance, is classified as a dry, not a fleshy fruit. Botanically, it is ''not'' a berry; it is an aggregate-accessory fruit, the latter term meaning the fleshy part is derived not from the plant's ovaries but from the receptacle that holds the
ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...
.Esau, K. (1977). ''Anatomy of seed plants''. John Wiley and Sons, New York. Numerous dry achenes are attached to the outside of the fruit-flesh, (see image); they appear to be seeds but each is actually an ovary of a flower, with a seed inside. Schizocarps are dry fruits, though some appear to be fleshy. They originate from syncarpous ovaries but do not actually dehisce; rather, they split into segments with one or more seeds. They include a number of different forms from a wide range of families, including carrot, parsnip, parsley, cumin.


Aggregate fruits

An aggregate fruit is also called an aggregation, or ''
etaerio A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
''; it develops from a single flower that presents numerous simple pistils (see graphic of raspberry pistils). Each pistil contains one carpel; together they form a fruitlet. The ultimate (fruiting) development of the aggregation of pistils is called an aggregate fruit, etaerio fruit, or simply an etaerio. Different types of aggregate fruits can produce different etaerios, such as achenes, drupelets, follicles, and berries. For example, the Ranunculaceae species, including '' Clematis'' and ''
Ranunculus ''Ranunculus'' is a large genus of about almost 1700 to more than 1800 species of flowering plants in the family Ranunculaceae. Members of the genus are known as buttercups, spearworts and water crowfoots. The genus is distributed in Europe ...
'', produces an etaerio of achenes; ''
Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species. Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of the ...
'' species, including raspberry: an etaerio of drupelets; ''
Calotropis ''Calotropis'' is a genus of flowering plants in the family Apocynaceae, first described as a genus in 1810. It is native to southern Asia and North Africa. They are commonly known as milkweeds because of the latex they produce. ''Calotropis'' ...
'' species: an etaerio of follicles fruit; '' Annona'' species: an etaerio of
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, ras ...
. Some other broadly recognized species and their etaerios (or aggregations) are: * Teasel; fruit is an aggregation of cypselas. * Tuliptree; fruit is an aggregation of samaras. *
Magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendr ...
and peony; fruit is an aggregation of follicles. *
American sweet gum American sweetgum (''Liquidambar styraciflua''), also known as American storax, hazel pine, bilsted, redgum, satin-walnut, star-leaved gum, alligatorwood, or simply sweetgum, is a deciduous tree in the genus ''Liquidambar'' native to warm temper ...
; fruit is an aggregation of capsules. * Sycamore; fruit is an aggregation of achenes. The '' raspberry''; its pistils are called ''drupelets'' because each pistil is like a small drupe attached to the receptacle. In some bramble fruits such as blackberry the receptacle, an accessory part, elongates and then develops as part of the fruit, making the blackberry an '' aggregate-accessory'' fruit. The strawberry is also an aggregate-accessory fruit, of which the seeds are contained in the achenes. Notably in all these examples, the fruit develops from a single flower, with numerous pistils.


Multiple fruits

A multiple fruit is formed from a cluster of flowers, (a 'multiple' of flowers) —also called an ''
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
''. Each ('smallish') flower produces a single fruitlet, which, as all develop, all merge into one mass of fruit. Examples include pineapple,
fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
, mulberry, Osage orange, breadfruit. An inflorescence (a cluster) of white flowers, called a head, is produced first. After fertilization, each flower in the cluster develops into a drupe; as the drupes expand, they develop as a '' connate'' organ, merging into a multiple fleshy fruit called a ''syncarp''. Progressive stages of multiple flowering and fruit development can be observed on a single branch of the Indian mulberry, or '' noni'', (see image). During the sequence of development, a progression of second, third, and more inflorescences are initiated in turn at the head of the branch or stem.


Accessory fruit forms

Fruits may incorporate tissues derived from other floral parts besides the ovary, including the receptacle, hypanthium, petals, or sepals. Accessory fruits occur in all three classes of fruit development—simple, aggregate, and multiple. Accessory fruits are frequently designated by the hyphenated term showing both characters. For example, a pineapple is a multiple-accessory fruit. a blackberry is an aggregate-accessory fruit, and an apple is a simple-accessory fruit.


Table of fleshy fruit examples


Seedless fruits

Seedlessness is an important feature of some fruits of commerce. Commercial cultivars of bananas and pineapples are examples of seedless fruits. Some cultivars of
citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...
fruits (especially grapefruit, mandarin oranges, navel oranges), satsumas, table grapes, and of watermelons are valued for their seedlessness. In some species, seedlessness is the result of ''
parthenocarpy In botany and horticulture, parthenocarpy is the natural or artificially induced production of fruit without fertilisation of ovules, which makes the fruit seedless. Stenospermocarpy may also produce apparently seedless fruit, but the seeds ar ...
'', where fruits set without fertilization. Parthenocarpic fruit-set may (or may not) require pollination, but most seedless citrus fruits require a stimulus from pollination to produce fruit. Seedless bananas and grapes are triploids, and seedlessness results from the abortion of the embryonic plant that is produced by fertilization, a phenomenon known as '' stenospermocarpy'', which requires normal pollination and fertilization.


Seed dissemination

Variations in fruit structures largely depend on the modes of dispersal applied to their seeds. Dispersal is achieved by wind or water, by explosive dehiscence, and by interactions with animals. Some fruits present their outer skins or shells coated with spikes or hooked burrs; these evolved either to deter would-be foragers from feeding on them, or to serve to attach themselves to the hair, feathers, legs, or clothing of animals, thereby using them as dispersal agents. These plants are termed
zoochorous In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...
; common examples include
cocklebur ''Xanthium'' (cocklebur) is a genus of flowering plants in the tribe Heliantheae within the family Asteraceae, native to the Americas and eastern Asia and some parts of south Asia . Description Cockleburs are coarse, herbaceous annual plants ...
,
unicorn plant ''Proboscidea'' is a genus of flowering plant in the family Martyniaceae, some of whose species are known as devil's claw, devil's horn, ram's horn, or unicorn plant. The plants produce long, hooked seed pods. The hooks catch on the feet of anim ...
, and beggarticks (or Spanish needle). By developments of mutual evolution the fleshy produce of fruits typically appeals to hungry animals, such that the seeds contained within are taken in, carried away and later deposited (i.e.,
defecated Defecation (or defaecation) follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus. The act has a variety of names ranging fro ...
) at a distance from the parent plant. Likewise, the nutritious, oily kernels of nuts typically motivate birds and squirrels to hoard them, burying them in soil to retrieve later during the winter of scarcity; thereby, uneaten seeds are sown effectively under natural conditions to germinate and grow a new plant some distance away from the parent. Other fruits have evolved flattened and elongated wings or helicopter-like blades, e.g., elm, maple, and tuliptree. This mechanism increases dispersal distance away from the parent via wind. Other wind-dispersed fruit have tiny " parachutes", e.g., dandelion,
milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to hum ...
, salsify. Coconut fruits can float thousands of miles in the ocean, thereby spreading their seeds. Other fruits that can disperse via water are
nipa palm ''Nypa fruticans'', commonly known as the nipa palm (or simply nipa, from ms, nipah) or mangrove palm, is a species of palm native to the coastlines and estuarine habitats of the Indian and Pacific Oceans. It is the only palm considered adap ...
and
screw pine ''Pandanus'' is a genus of monocots with some 750 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. The greatest number of species are found in Madagascar and Malaysia. Common names ...
. Some fruits have evolved propulsive mechanisms that fling seeds substantial distances —(perhaps up to 100 m in the case of the sandbox tree)— via explosive dehiscence or other such mechanisms, (see impatiens and
squirting cucumber ''Ecballium'' is a genus of flowering plants in the family Cucurbitaceae containing a single species, ''Ecballium elaterium'', also called the squirting cucumber or exploding cucumber (not the same plant as ''Cyclanthera brachystachya''). Its ...
.


Food uses

A cornucopia of fruits —fleshy (simple) fruits from apples to berries to watermelon; dry (simple) fruits including beans and rice, coconuts and carrots; aggregate fruits including strawberries, raspberries, blackberries, pawpaw; multiple fruits such as pineapple, fig, mulberries; (see above re all)— are commercially valuable as human food. They are eaten both fresh and as jams, marmalade and other fruit preserves. They are used extensively in manufactured and processed foods (cakes, cookies, baked goods, flavorings, ice cream, yogurt, canned vegetables, frozen vegetables and meals) and beverages such as fruit juices and alcoholic beverages (brandy, fruit beer, wine). Spices like vanilla, black pepper, paprika, and allspice are derived from berries. Olive fruit is pressed for olive oil and similar processing is applied to other oil bearing fruits/vegetables. Fruits are also used for socializing and gift-giving in the form of fruit baskets and fruit bouquets. Typically, many botanical fruits —"vegetables" in culinary ''parlance''— (including tomato, green beans, leaf greens, bell pepper, cucumber, eggplant, okra, pumpkin, squash, zucchini) are bought and sold daily in fresh produce markets and
greengrocer A greengrocer is a person who owns or operates a shop selling primarily fruit and vegetables. The term may also be used to refer to a shop selling primarily produce. It is used predominantly in the United Kingdom and Australia. In the Uni ...
ies and carried back to kitchens, at home or restaurant, for preparation of meals.


Storage

All fruits benefit from proper post harvest care, and in many fruits, the plant hormone ethylene causes ripening. Therefore, maintaining most fruits in an efficient cold chain is optimal for post harvest storage, with the aim of extending and ensuring shelf life.Why Cold Chain for Fruits:


Nutritional value

Various culinary fruits provide significant amounts of
fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
and water, and many are generally high in vitamin C. An overview of numerous studies showed that fruits (e.g., whole apples or whole oranges) are satisfying (filling) by simply eating and chewing them. The dietary fiber consumed in eating fruit promotes satiety, and may help to control body weight and aid reduction of blood
cholesterol Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol (or modified steroid), a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell memb ...
, a
risk factor In epidemiology, a risk factor or determinant is a variable associated with an increased risk of disease or infection. Due to a lack of harmonization across disciplines, determinant, in its more widely accepted scientific meaning, is often u ...
for cardiovascular diseases. Fruit consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. Regular consumption of fruit is generally associated with reduced risks of several diseases and functional declines associated with aging.


Food safety

For food safety, the
CDC The Centers for Disease Control and Prevention (CDC) is the national public health agency of the United States. It is a United States federal agency, under the Department of Health and Human Services, and is headquartered in Atlanta, Georg ...
recommends proper fruit handling and preparation to reduce the risk of food contamination and foodborne illness. Fresh fruits and vegetables should be carefully selected; at the store, they should not be damaged or bruised; and precut pieces should be refrigerated or surrounded by ice. All fruits and vegetables should be rinsed before eating. This recommendation also applies to produce with rinds or skins that are not eaten. It should be done just before preparing or eating to avoid premature spoilage. Fruits and vegetables should be kept separate from raw foods like meat, poultry, and seafood, as well as from utensils that have come in contact with raw foods. Fruits and vegetables that are not going to be cooked should be thrown away if they have touched raw meat, poultry, seafood, or eggs. All cut, peeled, or cooked fruits and vegetables should be refrigerated within two hours. After a certain time, harmful bacteria may grow on them and increase the risk of foodborne illness.


Allergies

Fruit allergies make up about 10 percent of all food related allergies.


Nonfood uses

Because fruits have been such a major part of the human diet, various cultures have developed many different uses for fruits they do not depend on for food. For example: *
Bayberry ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America and South America, and missing only from Aust ...
fruits provide a wax often used to make candles; * Many dry fruits are used as decorations or in dried flower arrangements (e.g.,
annual honesty ''Lunaria annua'', called honesty or annual honesty in English, is a species of flowering plant in the cabbage and mustard family Brassicaceae. It is native to the Balkans and south west Asia, and cultivated throughout the temperate world. Des ...
, cotoneaster, lotus,
milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to hum ...
,
unicorn plant ''Proboscidea'' is a genus of flowering plant in the family Martyniaceae, some of whose species are known as devil's claw, devil's horn, ram's horn, or unicorn plant. The plants produce long, hooked seed pods. The hooks catch on the feet of anim ...
, and wheat). Ornamental trees and shrubs are often cultivated for their colorful fruits, including
beautyberry ''Callicarpa'' (beautyberry) is a genus of shrubs and small trees in the family Lamiaceae.Heywood, V.H., Brummitt, R.K., Culham, A. & Seberg, O. 2007: Flowering Plant Families of the World. Royal Botanic Gardens, Kew. They are native to eas ...
, cotoneaster, holly, pyracantha, skimmia, and viburnum. * Fruits of opium poppy are the source of opium, which contains the drugs codeine and
morphine Morphine is a strong opiate that is found naturally in opium, a dark brown resin in poppies ('' Papaver somniferum''). It is mainly used as a pain medication, and is also commonly used recreationally, or to make other illicit opioids. T ...
, as well as the biologically inactive chemical theabaine from which the drug oxycodone is synthesized. * Osage orange fruits are used to repel cockroaches. * Many fruits provide
natural dye Natural dyes are dyes or colorants derived from plants, invertebrates, or minerals. The majority of natural dyes are vegetable dyes from plant sources— roots, berries, bark, leaves, and wood—and other biological sources such as fungi. Ar ...
s (e.g., cherry, mulberry, sumac, and walnut). * Dried gourds are used as bird houses, cups, decorations, dishes, musical instruments, and water jugs. * Pumpkins are carved into
Jack-o'-lantern A jack-o'-lantern (or jack o'lantern) is a carved lantern, most commonly made from a pumpkin or a root vegetable such as a rutabaga or turnip. Jack-o'-lanterns are associated with the Halloween holiday. Its name comes from the reported phen ...
s for Halloween. * The fibrous core of the mature and dry Luffa fruit is used as a sponge. * The spiny fruit of burdock or
cocklebur ''Xanthium'' (cocklebur) is a genus of flowering plants in the tribe Heliantheae within the family Asteraceae, native to the Americas and eastern Asia and some parts of south Asia . Description Cockleburs are coarse, herbaceous annual plants ...
inspired the invention of Velcro. * Coir fiber from coconut shells is used for brushes, doormats, floor tiles, insulation, mattresses, sacking, and as a growing medium for container plants. The shell of the coconut fruit is used to make bird houses, bowls, cups, musical instruments, and souvenir heads. * The hard and colorful grain fruits of Job's tears are used as decorative beads for jewelry, garments, and ritual objects. * Fruit is often a subject of still life paintings.


See also

* Fruit tree *
Fruitarianism Fruitarianism () is a diet related to veganism that consists primarily of consuming fruits and possibly nuts and seeds, but without any animal products. Fruitarian diets are subject to criticism and health concerns. Fruitarianism may be adopted ...
* List of countries by fruit production * List of culinary fruits * List of foods * List of fruit dishes


References


Further reading

* Gollner, Adam J. (2010). ''The Fruit Hunters: A Story of Nature, Adventure, Commerce, and Obsession''. Scribner. * Watson, R. R., and Preedy, V.R. (2010, eds.). ''Bioactive Foods in Promoting Health: Fruits and Vegetables''. Academic Press.


External links


Images of fruit development from flowers
at bioimages.Vanderbilt.edu

at bioimages.Vanderbilt.edu

from California Rare Fruit Growers, Inc.
Photo ID of Fruits
by Capt. Pawanexh Kohli * {{Authority control Fruit Pollination Types of food