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A flyway is a
flight path In the United States, airways or air routes are defined by the Federal Aviation Administration (FAA) in two ways: "VOR Federal airways and Low/Medium Frequency (L/MF) (Colored) Federal airways" These are designated routes which aeroplanes f ...
used by large numbers of birds while migrating between their breeding grounds and their
overwintering Overwintering is the process by which some organisms pass through or wait out the winter season, or pass through that period of the year when "winter" conditions (cold or sub-zero temperatures, ice, snow, limited food supplies) make normal act ...
quarters. Flyways generally span
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
s and often pass over
ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wor ...
s. Although applying to any species of migrating bird, the concept was first conceived and applied to
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which ...
and shore birds. The flyways can be thought of as wide arterial highways to which the migratory routes of different species are
tributaries A tributary, or affluent, is a stream or river that flows into a larger stream or main stem (or parent) river or a lake. A tributary does not flow directly into a sea or ocean. Tributaries and the main stem river drain the surrounding drain ...
. An alternative definition is that a flyway is the entire range of a migratory bird, encompassing both its breeding and non-breeding grounds, and the resting and feeding locations it uses while migrating. There are four major north–south flyways in North America and six covering Eurasia, Africa, and Australasia.


History

The passing of the
Migratory Bird Treaty Act of 1918 The Migratory Bird Treaty Act of 1918 (MBTA), codified at (although §709 is omitted), is a United States federal law, first enacted in 1918 to implement the convention for the protection of migratory birds between the United States and Canada ...
in the United States resulted in a need for more information on bird migration.
Frederick Charles Lincoln Frederick Charles Lincoln (5 May 1892 – 16 September 1960) was an American ornithologist. Early life and family Lincoln was born on 5 May 1892 in Denver, Colorado. Career As a teenager working at the Colorado Museum of Natural History in 1 ...
was put in charge and improved methods for trapping and banding, developed record-keeping procedures, recruited banders, fostered international cooperation, and promoted banding as a tool for research and wildlife management. He found it was possible to establish the routes used by waterfowl during their annual migrations and he developed the flyway concept, a key idea in the management and regulation of hunting of migratory birds; by establishing the routes used, estimates of population sizes could be made and suitable protection could be put in place. The special vulnerability of waterfowl and shorebirds on their international migrations, with their specific needs for suitable wetland stopovers, resulted in the signing of the
Ramsar Convention The Ramsar Convention on Wetlands of International Importance Especially as Waterfowl Habitat is an international treaty for the conservation and sustainable use of Ramsar sites (wetlands). It is also known as the Convention on Wetlands. It ...
in 1971. As a result, over 2300
Ramsar sites A Ramsar site is a wetland site designated to be of international importance under the Ramsar Convention,8 ha (O) *** Permanent 8 ha (P) *** Seasonal Intermittent < 8 ha(Ts) ** Marshes on inorganic soils: *** Permanent (herb dominated) (Tp) *** P ...
have been established around the world, many being situated on flyways where they provide the vital habitat needed by the birds on their journeys.


Flyways

Many bird populations migrate long distances twice a year. The most common pattern involves flying north in the spring to breed in the temperate parts of the northern hemisphere or the
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
during summer and returning southward in the autumn to wintering grounds in warmer regions, often on the other side of the equator. A similar pattern occurs in the southern hemisphere with birds flying south to breed and north to overwinter, but on a much smaller scale. The flyway, or route, taken by different bird species varies, but each population has its traditional staging points along the route where birds feed to build up their energy reserves to prepare for the next migratory stage; the route used on the spring migration may be different from that used in the autumn and will depend on such factors as wind direction and the availability of food at staging points. Flyways may not be the shortest route available but may have curves or doglegs. Birds of different species may follow similar routes, and populations from one area may merge with other groups and diverge to reach different destinations. Flyways tend to avoid obstacles such as mountain ranges and oceans, running parallel to the barriers and following routes along the coast or along major river valleys. Passerines often fly on a broader front across the terrain, either flying over or circumventing obstacles on the route, according to their evolutionary adaptations. In selecting routes, birds may overcompensate for predicted winds. Terrestrial birds tend to travel over land, raptors need routes where
thermal A thermal column (or thermal) is a rising mass of buoyant air, a convective current in the atmosphere, that transfers heat energy vertically. Thermals are created by the uneven heating of Earth's surface from solar radiation, and are an example ...
s can give them the lift they require, sea birds prefer ocean routes and wetland birds need routes with suitable staging sites; deltas and coastal wetlands provide reliable food sources for this purpose whereas inland wetlands are less predictable.


Flyways of the Americas

The Atlantic Flyway starts in northern Canada and
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
and follows the Atlantic coast of Canada and the United States to the
Caribbean Sea The Caribbean Sea ( es, Mar Caribe; french: Mer des Caraïbes; ht, Lanmè Karayib; jam, Kiaribiyan Sii; nl, Caraïbische Zee; pap, Laman Karibe) is a sea of the Atlantic Ocean in the tropics of the Western Hemisphere. It is bounded by Mexic ...
, and on to tropical South America. The
Mississippi Flyway The Mississippi Flyway is a bird migration route that generally follows the Mississippi, Missouri, and Lower Ohio Rivers in the United States across the western Great Lakes to the Mackenzie River and Hudson Bay in Canada. The main endpoint ...
starts from northeastern Canada and passes over the Great Lakes, following the lower
Ohio River The Ohio River is a long river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing southwesterly from western Pennsylvania to its mouth on the Mississippi River at the southern tip of ...
, the
Missouri Missouri is a state in the Midwestern region of the United States. Ranking 21st in land area, it is bordered by eight states (tied for the most with Tennessee): Iowa to the north, Illinois, Kentucky and Tennessee to the east, Arkansas t ...
and the
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
to the
Gulf of Mexico The Gulf of Mexico ( es, Golfo de México) is an ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of the United ...
, and on to Central and South America. There are no mountain barriers on this route. The
Central Flyway The Central Flyway is a bird migration route that generally follows the Great Plains in the United States and Canada. The main endpoints of the flyway include the Canadian Prairies and the region surrounding the Gulf of Mexico; the migration ...
starts from central Canada and crosses the
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, a ...
before continuing southwards to the Gulf of Mexico, merging with the Mississippi Flyway. There are no mountain barriers on this route. The
Pacific Flyway The Pacific Flyway is a major north-south flyway for migratory birds in the Americas, extending from Alaska to Patagonia. Every year, migratory birds travel some or all of this distance both in spring and in fall, following food sources, heading ...
is a north–south flyway for birds migrating from breeding grounds in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
and Canada to their overwintering areas in South America, some species travelling as far south as
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
. The Allegheny Front flyway in the central
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. The ...
is an important flyway for migratory birds traveling from their northern breeding grounds to their southern wintering sites.


Flyways of Eurasia, Africa, and Australasia

The
East Atlantic Flyway The East Atlantic Flyway is a migration route used by about 90 million birds annually, passing from their breeding areas in the United States, Canada, Greenland, Iceland, Siberia and northern Europe to wintering areas in western Europe and on t ...
starts from northern North America,
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
,
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
, northern Europe and western Siberia and leads to wintering areas in western Europe and North Africa, with some birds continuing down the west coast of the continent to South Africa. The Black Sea-Mediterranean Flyway starts from northern and western Siberia and leads across Asia, the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Rom ...
and the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
to northern Africa. Little has been published about birds using this flyway. The Asian–East African Flyway starts from the northern breeding grounds of water birds in Siberia and leads across Asia to East Africa. Little has been published about birds using this flyway. The Central Asian-Indian Flyway starts from the northern breeding grounds of water birds in Siberia and leads across Asia to the Indian subcontinent. Little has been published about birds using this flyway. The East Asian–Australasian Flyway starts at the
Taymyr Peninsula The Taymyr Peninsula (russian: Таймырский полуостров, Taymyrsky poluostrov) is a peninsula in the Far North of Russia, in the Siberian Federal District, that forms the northernmost part of the mainland of Eurasia. Administrat ...
in Russia and
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
and extends southwards to southeastern Asia, Australia and New Zealand. This flyway overlaps with the West Pacific Flyway. About 60 species of shorebird use this route. The West Pacific Flyway links New Zealand and the east coast of Australia, through the central Pacific Ocean and the east coast of northern Asia, including Japan and the Korean Peninsula, ending up in eastern Siberia, including the Chukchi and
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
peninsulas, and Alaska. This flyway overlaps with the East Asian–Australasian Flyway.


References

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