In terrestrial

vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...

s, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the ground, and the rest of its foot lifted. Digitigrades include walking birds (what many assume to be bird knees are actually

ankle The ankle, or the talocrural region, or the jumping bone (informal) is the area where the foot and the leg meet. The ankle includes three joints: the ankle joint proper or talocrural joint, the subtalar joint, and the inferior tibiofibular join ...

s), cats, dogs, and many other

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s, but not

plantigrade 151px, Portion of a human skeleton, showing plantigrade habit In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...

s or

unguligrade Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, ca ...

s. Digitigrades generally move more quickly and quietly than other animals. Les différents types de locomotions chez les amniotes

There are anatomical differences between the limbs of plantigrades, like humans, and both unguligrade and digitigrade limbs. Digitigrade and unguligrade animals have relatively long

carpal The carpal bones are the eight small bones that make up the wrist (or carpus) that connects the hand to the forearm. The term "carpus" is derived from the Latin carpus and the Greek καρπός (karpós), meaning "wrist". In human anatomy, the ...

s and tarsals, and the bones which would correspond to the human ankle are thus set much higher in the limb than in a human. In a digitigrade animal, this effectively lengthens the foot, so much so that what are often thought of as a digitigrade animal's "hands" and "feet" correspond only to what would be the

bones A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, an ...

of the human finger or toe. Humans usually walk with the soles of their feet on the ground, in plantigrade locomotion. In contrast, digitigrade animals walk on their

distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

and

intermediate phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...

. Digitigrade locomotion is responsible for the distinctive hooked shape of dog legs. Unguligrade animals, such as horses and cattle, walk only on the distal-most tips of their digits, while in digitigrade animals, more than one segment of the digit makes contact with the ground, either directly (as in birds) or via paw-pads (as in dogs).

Examples

Mesonychids Mesonychia ("middle claws") is an extinct taxon of small- to large-sized carnivorous ungulates related to artiodactyls. Mesonychids first appeared in the early Paleocene, went into a sharp decline at the end of the Eocene, and died out entirely ...

Dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

(digitigrade and semi-digitigrade) **

Birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

(except for

loons Loons ( North American English) or divers (British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order ...

and grebes which are plantigrade) *

Pigs The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus s ...

(semi-digitigrade) *

Hippos A hippo or hippopotamus is either of two species of large African mammal which live mainly in and near water: * Hippopotamus * Pygmy hippopotamus Hippo or Hippos may also refer to: Toponymy * The ancient city of Hippo Regius (modern Annaba, Al ...

(semi-digitigrade) * ''

Pakicetus ''Pakicetus'' is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to Pakistan during the Eocene, about 50 million years ago. It was a wolf-like animal, about to long, and lived in and around water where it a ...

'' * ''

Indohyus ''Indohyus'' is an extinct genus of digitigrade artiodactyls known from Eocene fossils in Asia. This small chevrotain-like animal found in the Himalayas is one of the earliest known non-cetacean ancestors of whales. Discovery The fossils were di ...

'' *

Thylacine The thylacine ( , or , also ) (''Thylacinus cynocephalus'') is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The last known live animal was captured in 1930 in Tasman ...

Cats The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of t ...

Hyena Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the clas ...

s *

Mongooses A mongoose is a small terrestrial carnivorous mammal belonging to the family Herpestidae. This family is currently split into two subfamilies, the Herpestinae and the Mungotinae. The Herpestinae comprises 23 living species that are native to ...

Canids Canidae (; from Latin, ''canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within ...

Elephants Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae and ...

(semi-digitigrade)APPEARANCE/ MORPHOLOGY: LEGS, SPINE AND TRACKS with literature reports for the Asian Elephant - Elephas maximus (editorial comment)
/ref> *

Capybaras The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydro ...

(semi-digitigrade) * ''

Trucidocynodon ''Trucidocynodon'' is an extinct genus of ecteniniid cynodonts from Upper Triassic of Brazil. It contains a single species, ''Trucidocynodon riograndensis''. Fossils of ''Trucidocynodon'' were discovered in Santa Maria Formation outcrops in Pal ...

'' (semi-digitigrade at least in the forelimbs)

References

External links

Yes, the Shin Bone Is Connected to the Ankle Bone
{{locomotion Terrestrial locomotion