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The corsac fox (''Vulpes corsac''), also known simply as a corsac, is a medium-sized fox found in
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate gras ...
s,
semi-desert A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi ...
s and
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s in
Central Asia Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the fo ...
, ranging into
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
and northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. Since 2004, it has been classified as ''
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
'' by
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, but populations fluctuate significantly, and numbers can drop tenfold within a single year. It is also known as the steppe fox. The word "corsac" is derived from the Russian name for the animal, ''"korsák"'' (корса́к), derived ultimately from Turkic "karsak".


Description

The Corsac fox is a medium-sized fox, with a head and body length of , and a tail long. Adults weigh from . It has grey to yellowish fur over much of the body, with paler underparts and pale markings on the mouth, chin, and throat. During the winter, the coat becomes much thicker and silkier in texture, and is straw-grey in colour, with a darker line running down the back. For a fox, it has small teeth and a wide skull. One source claims that this species can climb trees and has been domesticated in the past. It is reported to have keen eyesight and hearing and an acute sense of smell. It has a number of
scent gland Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and s ...
s, some of which produce pungent odors, although not so extreme as those found in some other ''Vulpes'' species. The glands are found in the anal region, above the base of the tail, and on the paws and cheeks. Corsac foxes are reported to bark during hunting or when threatening rivals, and to use higher pitch yelps or chirps as alarm calls or social greetings.


Distribution and habitat

Corsac foxes live in the
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate gras ...
s and semidesert of central and northeast
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
. They are found throughout
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...
,
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked co ...
, and
Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the s ...
, and through all except the northernmost regions of
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
. In the south, their range extends into the more northern parts of
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...
,
Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the ea ...
,
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is borde ...
, and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
, and they can also be found in neighbouring regions of
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
. Three subspecies are currently recognised: * ''Vulpes corsac corsac'' - northern Kazakhstan, southern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part ...
* ''V. c. kalmykorum'' - northern Uzbekistan,
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
* ''V. c. turkmenicus'' - southern Uzbekistan, Turkmenistan, China, Mongolia, and neighbouring regions These foxes inhabit open grassy steppes and semideserts, and avoid dense vegetation and mountainous regions. True deserts with drifting sands are also avoided, as are snowfields more than about deep. Corsac foxes generally stay far away from human disturbances.


Ecology and behavior

As an adaption to the arid climate in which they live, corsac foxes can forego food and water for extended periods of time. The corsac fox is an opportunistic forager and hunter. Its diet varies throughout its range, but consists foremost of small and medium-sized vertebrates, insects and small rodents, such as
vole Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s,
gerbil The Mongolian gerbil or Mongolian jird (''Meriones unguiculatus'') is a small rodent belonging to the subfamily Gerbillinae. Their body size is typically , with a tail, and body weight , with adult males larger than females. The animal is us ...
s,
jerboa Jerboas (from ar, جربوع ') are hopping desert rodents found throughout North Africa and Asia, and are members of the family Dipodidae. They tend to live in hot deserts. When chased, jerboas can run at up to . Some species are preyed on b ...
s,
hamster Hamsters are rodents (order Rodentia) belonging to the subfamily Cricetinae, which contains 19 species classified in seven genera.Fox, Sue. 2006. ''Hamsters''. T.F.H. Publications Inc. They have become established as popular small pets. The ...
s, and ground squirrels. It also feeds opportunistically on larger prey including
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The g ...
s and pikas. It scavenges for
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
and human refuse as well. Although predominantly carnivorous, it occasionally eats fruit and other vegetation, especially during winter when availability of animal prey is low. Natural predators of the corsac fox include
gray wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly un ...
,
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and femal ...
,
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, j ...
s, and (rarely, for pups) the upland buzzard.
Golden eagles The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds ...
are major predators, with 38% of remains found belonging to the corsac fox at a golden eagle nest in Mongolia.Ellis, D.H., Tsengeg, P., Whitlock, P. & Ellis, M.H. 2000. ''Predators as prey at a Golden Eagle ''Aquila chrysaetos'' eyrie in Mongolia''. Ibis, 142: 139 – 141. The Corsac fox is a nocturnal and nomadic hunter of the steppes. It does not have a defended
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
, and unlike some foxes, sometimes forms packs. Diurnal activity is more common at times when kits need to be fed and when food is scarce, as during the winter. Because it cannot hunt in deep snow, it either shelters in dens during harsh weather, or, in the northern parts of its range, it migrates up to south in the winter. It has been reported to follow herds of local antelope, relying on them to compress the snow as they pass. It digs its own dens, which are generally shallow, but also takes over the burrows of other animals, such as marmots, ground squirrels, or
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by ...
s. Dens may have several entrances, but are usually less than deep. The burrow is shared between the social packs, with several dens and connecting holes, which resemble "corsac cities". They are excellent climbers, but are rather slow runners and could be caught easily by a dog. While they are reported to be
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
in the wild, in captivity they are very active during the day.


Reproduction

The mating season starts in January and ends in March. Males will initially fight for access to females, but eventually establish a
monogamous Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time ( serial monogamy) — as compared to the various forms of non-monogamy (e.g., pol ...
bond, and assist in the raising of their young. The mother initially creates a birthing den, which is sometimes shared with other pregnant females, but moves her young to new burrows several times after they are born. Typically, two to six young are born after a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pr ...
period of 52 to 60 days, although cases of ten kits being born in a single litter have been reported. Newborn kits weigh around , and have fluffy, light brown fur that turns yellowish as they age. They are born blind, and open their eyes at around two weeks of age; they begin to eat meat at four weeks, and emerge from the den shortly after. Corsac foxes reach
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definit ...
within 9 to 10 months and reproduce in the second year of life. They live up to 9 years in the wild.


Evolution

The corsac fox is one species within a
holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
of foxes that also includes the
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
, the
swift fox The swift fox (''Vulpes velox'') is a small light orange-tan fox around the size of a domestic cat found in the western grasslands of North America, such as Montana, Colorado, New Mexico, Kansas, Oklahoma and Texas. It also lives in southern M ...
and the
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in ...
, all of which it resembles. However, the closest related species to the corsac fox is probably the Tibetan sand fox. The immediate ancestor of the corsac fox is believed to be the
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
''Vulpes praecorsac'', which lived in central Europe during the early
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
. Fossils of corsac foxes date back to the mid-Pleistocene, and show the species once reached as far west as Switzerland, and as far south as
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a p ...
. Very recent subfossil remains dating to 3000 to 1000 B.C.E. have in found in the
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a p ...
and 0 to 1500 B.C.E. in the Donev River Basin in the northern
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Rom ...
region.


Threats

The major threat posed to the corsac fox is poaching, as it is a valuable fur-bearing species and has been harvested by humans since the
Bronze Age The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
for subsistence and commercial purposes. It is a slow runner and therefore easily caught by hunters; the population has been reduced in areas where it has been heavily hunted for its fur. Traditionally, the corsac fox is often hunted with trained domestic dogs, saker falcons, and
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird ...
s, as well as caught with traps set at burrow entrances, leghold traps, and firearms. In the late 19th century, up to 10,000 foxes were killed annually for pelt trade. The general population remains healthy, however, as the corsac fox has proven to be able to withstand great hunting pressures, and their habitats remain intact due to the low human population density in its range. Other threats include overgrazing by livestock and landscape development; the decline of marmots may also impact the species in some areas, as it often uses marmot burrows as daytime resting locations. The other main threat is natural disasters, which can cause the numbers of foxes to drop 90% in some areas, but the population often recovers quickly. , the corsac fox is listed as
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. ...
in the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolo ...
.


References


External links

* {{Taxonbar, from=Q187024 corsac fox Mammals of Afghanistan Mammals of Mongolia Mammals of China Mammals of Central Asia corsac fox Taxa named by Carl Linnaeus