Climax species, also called late

seral A seral community is an intermediate stage found in ecological succession in an ecosystem advancing towards its climax community. In many cases more than one seral stage evolves until climax conditions are attained. A prisere is a collection of se ...

, late-successional,

K-selected In ecology, ''r''/''K'' selection theory relates to the selection of combinations of traits in an organism that trade off between quantity and quality of offspring. The focus on either an increased quantity of offspring at the expense of indivi ...

or equilibrium species, are

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

that can germinate and grow with limited resources; e.g., they need heat exposure or low water availability. They are the species within forest succession that are more adapted to stable and predictable environments, and will remain essentially unchanged in terms of species composition for as long as a site remains undisturbed. The

seedlings A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embryo ...

of climax species can grow in the shade of the parent trees, ensuring their dominance indefinitely. The presence of climax species can also reduce the prevalence of other species within an ecosystem. However, a disturbance, such as fire, may kill the climax species, allowing

pioneer Pioneer commonly refers to a settler who migrates to previously uninhabited or sparsely inhabited land. In the United States pioneer commonly refers to an American pioneer, a person in American history who migrated west to join in settling and de ...

or earlier successional species to re-establish for a time. They are the opposite of

pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so ...

, also known as

ruderal A ruderal species is a plant species that is first to colonize disturbed lands. The disturbance may be natural for example, wildfires or avalanchesor the consequences of human activities, such as construction ( of roads, of buildings, mining, et ...

, fugitive, opportunistic or

R-selected In ecology, ''r''/''K'' selection theory relates to the selection of combinations of traits in an organism that trade off between quantity and quality of offspring. The focus on either an increased quantity of offspring at the expense of individ ...

species, in the sense that climax species are good competitors but poor colonizers, whereas pioneer species are good colonizers but poor competitors. Given the prevailing ecological conditions, climax species dominate the

climax community In scientific ecology, climax community or climatic climax community is a historic term for a community of plants, animals, and fungi which, through the process of ecological succession in the development of vegetation in an area over time, hav ...

. When the pace of succession slows down as the result of ecological homeostasis, the maximum permitted

biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...

is reached. Their reproductive strategies and other adaptive characteristics can be considered more sophisticated than those of opportunistic species. Through negative feedback, they adapt themselves to specific environmental conditions. Climax species are mostly found in forests. Climax species, closely controlled by carrying capacity, follow K strategies, wherein species produce fewer numbers of potential offspring, but invest more heavily in securing the reproductive success of each one to the micro-environmental conditions of its specific

ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...

. Climax species might be

iteroparous Semelparity and iteroparity are two contrasting reproductive strategies available to living organisms. A species is considered semelparous if it is characterized by a single reproductive episode before death, and iteroparous if it is characteri ...

, energy consumption efficient and

nutrient cycling A nutrient cycle (or ecological recycling) is the movement and exchange of inorganic and organic matter back into the production of matter. Energy flow is a unidirectional and noncyclic pathway, whereas the movement of mineral nutrients is cycli ...

Disputed term

The idea of a climax species has been criticized in recent ecological literature. Any assessment of successional states depends on assumptions about the natural fire regime. But the idea of a dominant species is still widely used in silvicultural programs and

California Department of Forestry The California Department of Forestry and Fire Protection (CAL FIRE) is the fire department of the California Natural Resources Agency in the U.S. state of California. It is responsible for fire protection in various areas under state responsib ...

literature.

Examples

White spruce White spruce is a common name for several species of spruce ('' Picea'') and may refer to: * ''Picea glauca'', native to most of Canada and Alaska with limited populations in the northeastern United States * '' Picea engelmannii'', native to the ...

(''Picea glauca'') is an example of a climax species in the northern forests of North America due to its ability to adapt to resource scarce, stable conditions, it dominates Northern forest ecosystem in the absence of a disturbance. Other examples of climax species in old-growth forests: * Canadian hemlock *

Pacific silver fir ''Abies amabilis'', commonly known as the Pacific silver fir, is a fir native to the Pacific Northwest of North America, occurring in the Pacific Coast Ranges and the Cascade Range. It is also commonly referred to as the white fir, red fir, l ...

White fir ''Abies concolor'', the white fir, is a coniferous tree in the pine family Pinaceae. This tree is native to the mountains of western North America, including the Cascade Range and southern Rocky Mountains, and into the isolated mountain ranges ...

Yellow carabeen ''Sloanea woollsii'', commonly known as yellow carabeen, is a large tree species with plank buttresses that is native to northeastern NSW and eastern Queensland, Australia. Its southern distributional limit is near the town of Bulahdelah (32� ...

Blue grama ''Bouteloua gracilis'', the blue grama, is a long-lived, warm-season ( C4) perennial grass, native to North America. It is most commonly found from Alberta, Canada, east to Manitoba and south across the Rocky Mountains, Great Plains, and U.S ...

* Douglas fir *

Coast redwood ''Sequoia sempervirens'' ()''Sunset Western Garden Book,'' 1995:606–607 is the sole living species of the genus '' Sequoia'' in the cypress family Cupressaceae (formerly treated in Taxodiaceae). Common names include coast redwood, coastal ...

European beech ''Fagus sylvatica'', the European beech or common beech is a deciduous tree belonging to the beech family Fagaceae. Description ''Fagus sylvatica'' is a large tree, capable of reaching heights of up to tall and trunk diameter, though more ...

Disputed term

Examples

See also

References

Further reading