The archerfish (spinner fish or archer fish) form a

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

, Toxotidae, of

fish Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...

known for their habit of preying on land-based

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...

s and other small animals by shooting them down with water droplets from their specialized mouths. The family is small, consisting of ten species in a single genus, ''Toxotes''. Most species live in fresh water rivers, streams and pools, but two or three are

euryhaline Euryhaline organisms are able to adapt to a wide range of salinities. An example of a euryhaline fish is the molly (''Poecilia sphenops'') which can live in fresh water, brackish water, or salt water. The green crab ('' Carcinus maenas'') is an ...

, inhabiting both fresh and

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estua ...

water habitats such as

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...

and

mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...

s. They can be found from India,

Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mo ...

and

Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...

, through

Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...

, to

Northern Australia The unofficial geographic term Northern Australia includes those parts of Queensland and Western Australia north of latitude 26° and all of the Northern Territory. Those local government areas of Western Australia and Queensland that lie p ...

and

Melanesia Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from Indonesia's New Guinea in the west to Fiji in the east, and includes the Arafura Sea. The region includes the four independent countries of Fiji, V ...

. Archerfish or spinnerfish bodies are deep and laterally compressed, with the

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...

, and the profile a straight line from dorsal fin to mouth. The mouth is protractile, and the lower jaw juts out. Sizes are fairly small, typically up to about , but ''T. chatareus'' can reach . Archerfish are popular for aquaria, but difficult to feed since they prefer live prey.

Capture of prey

Archerfish are remarkably accurate in their

shooting Shooting is the act or process of discharging a projectile from a ranged weapon (such as a gun, bow, crossbow, slingshot, or blowpipe). Even the acts of launching flame, artillery, darts, harpoons, grenades, rockets, and guided missiles ...

; an adult fish almost always hits the target on the first shot. Although it is presumed that all archerfish species do this, it has only been confirmed from ''T. blythii'', ''T. chatareus'' and ''T. jaculatrix''. They can bring down insects and other prey up to above the water's surface. This is partially due to their good eyesight, but also to their ability to compensate for the

refraction In physics, refraction is the redirection of a wave as it passes from one medium to another. The redirection can be caused by the wave's change in speed or by a change in the medium. Refraction of light is the most commonly observed phenomen ...

of light as it passes through the air-water interface when aiming at their prey. They typically spit at prey at a mean angle of about 74° from the horizontal but can still aim accurately when spitting at angles between 45° and 110°. When an archerfish selects its prey, it rotates its eye so that the image of the prey falls on a particular portion of the eye in the ventral temporal periphery of the retina, and its lips just break the surface, squirting a jet of water at its victim. The archerfish does this by forming a small groove in the roof of its mouth and its tongue into a narrow channel. It then fires by contracting its gill covers and forcing water through the channel, shooting a stream that, shaped by its mouth parts, travels faster at the rear than at the front. This speed differential causes the stream to become a blob directly before impact as the slower leading water is overtaken by the faster trailing water, and it is varied by the fish to account for differences in range. It also makes this one of the few animals that both make and use

tools A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ba ...

, as they both utilise the water and shape it to make it more useful to them. They are persistent and will make multiple shots if the first one fails. Young archerfish start shooting when they are about long but are inaccurate at first and must learn from experience. During this learning period, they hunt in small

schools A school is an educational institution designed to provide learning spaces and learning environments for the teaching of students under the direction of teachers. Most countries have systems of formal education, which is sometimes compulsor ...

. This way, the probability is enhanced that at least one jet will hit its target. A 2006 experimental study found that archerfish appear to benefit from

observational learning Observational learning is learning that occurs through observing the behavior of others. It is a form of social learning which takes various forms, based on various processes. In humans, this form of learning seems to not need reinforcement to o ...

by watching a performing group member shoot, without having to practice: However, little of their social behaviour is currently known beyond that archerfish are sensitive to, and make changes to their shooting behaviour, when conspecifics are visible to them. This is probably as a result of the potential threat of

kleptoparasitism Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when ...

that other archerfish represent to a shooting fish. An archerfish will often leap out of the water and grab an insect in its mouth if it happens to be within reach. Individuals typically prefer to remain close to the surface of the water. New research has found that archerfish also use jets to hunt underwater prey, such as those embedded in silt. It is not known whether they learned aerial or underwater shooting first, but the two techniques may have evolved in parallel, as improvements in one can be adapted to the other.

Species

There are 9 valid species, 8 in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

''Toxotes'': *'' Protoxotes lorentzi''

Weber Weber (, or ; German: ) is a surname of German origin, derived from the noun meaning " weaver". In some cases, following migration to English-speaking countries, it has been anglicised to the English surname 'Webber' or even 'Weaver'. Notable pe ...

, 1910 - primitive archerfish *''

Toxotes blythii ''Toxotes blythii'', the clouded archerfish or zebra archerfish, is a perciform fish of genus ''Toxotes''. It is found in rivers and estuaries in Myanmar, ranging from the lower Irrawaddy to the Tenasserim Division, including lower Sittaung and ...

Boulenger Boulenger is a surname. Notable people with the surname include: * Benjamin Boulenger (born 1990), French footballer * Edward George Boulenger (1888–1946), British zoologist, director of aquarium at London Zoo * George Albert Boulenger (1858– ...

, 1892 - clouded archerfish, zebra archerfish *''Toxotes carpentariensis'', Castelnau, 1878 *''

Toxotes chatareus ''Toxotes chatareus'', sometimes known by the common names common archerfish, seven-spot archerfish or largescale archerfish, is a species of perciform fish in the archerfish genus '' Toxotes''.Martin F. Gomon & Dianne J. Bray (2011Sevenspot Arc ...

'' (

Hamilton Hamilton may refer to: People * Hamilton (name), a common British surname and occasional given name, usually of Scottish origin, including a list of persons with the surname ** The Duke of Hamilton, the premier peer of Scotland ** Lord Hamilto ...

, 1822) - largescale archerfish, common archerfish *'' Toxotes jaculatrix'' (

Pallas Pallas may refer to: Astronomy * 2 Pallas asteroid ** Pallas family, a group of asteroids that includes 2 Pallas * Pallas (crater), a crater on Earth's moon Mythology * Pallas (Giant), a son of Uranus and Gaia, killed and flayed by Athena * Pa ...

, 1767) - banded archerfish *'' Toxotes kimberleyensis''

Allen Allen, Allen's or Allens may refer to: Buildings * Allen Arena, an indoor arena at Lipscomb University in Nashville, Tennessee * Allen Center, a skyscraper complex in downtown Houston, Texas * Allen Fieldhouse, an indoor sports arena on the Univer ...

, 2004 - Kimberley archerfish, western archerfish *''

Toxotes microlepis The smallscale archerfish (''Toxotes microlepis'') is a perciform fish of genus ''Toxotes''. As its name suggests, the scales of the smallscale archerfish are smaller than those of other archerfish. They reach a maximum length of . Smallscale ar ...

'' Günther, 1860 - smallscale archerfish *'' Toxotes oligolepis'' Bleeker, 1876 - big scale archerfish *''

Toxotes sundaicus Toxotes may refer to: Biology * Banded archerfish (''Toxotes jaculatrix''), Pallas 1767 * ''Toxotes chatareus'', Hamilton 1822 * Smallscale archerfish The smallscale archerfish (''Toxotes microlepis'') is a perciform fish of genus ''Toxotes''. ...

Kottelat Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandCo ...

Tan Tan or TAN may refer to: Businesses and organisations * Black and Tans, a nickname for British special constables during the Irish War of Independence. By extension "Tans" can now also colloquially refer to English or British people in general, es ...

, 2018

Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pala ...

from: -55.8 till: -33.9 color:eocene text:

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', ...

from: -33.9 till: -23.03 color:oligocene text:

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but t ...

from: -23.03 till: -5.332 color:miocene text:

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:

Paleogene The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning o ...

from: -23.03 till: -2.588 color:neogene text:

Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...

from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:eocene bar:NAM1 from: -55.8 till: 0 text:

Toxotes Toxotes may refer to: Biology * Banded archerfish (''Toxotes jaculatrix''), Pallas 1767 * ''Toxotes chatareus'', Hamilton 1822 * Smallscale archerfish (''Toxotes microlepis''), Günther 1860 * Big scale archerfish (''Toxotes oligolepis''), Bleeker ...

PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

from: -33.9 till: -23.03 color:oligocene text:

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text: Pleist. from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q.

References

External links

Archerfish Shooting Various Insects

– LiveScience.com {{Taxonbar, from=Q732534 Monogeneric fish families Articles containing video clips Taxa named by Georges Cuvier

Capture of prey

Species

Timeline

See also

References

External links