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Fur is a thick growth of
hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...
that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket that keeps the animal warm. The fur of mammals has many uses: protection, sensory purposes, waterproofing, and camouflaging, with the primary usage being thermoregulation. The types of hair include * ''definitive'', which may be
shed A shed is typically a simple, single-story roofed structure that is used for hobbies, or as a workshop in a back garden or on an allotment. Sheds vary considerably in their size and complexity of construction, from simple open-sided ones de ...
after reaching a certain length; * ''
vibrissae Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarse ...
'', which are sensory hairs and are most commonly whiskers; * ''pelage'', which consists of guard hairs, under-fur, and
awn hair Awn hairs are the intermediate hairs in a mammal's coat. They are shorter than the guard hairs and longer than the down hairs. They help with insulation and protect the down hairs underneath. Most of the visible coat is made of this kind of hair. ...
; * '' spines'', which are a type of stiff guard hair used for defense in, for example,
porcupine Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predation. The term covers two families of animals: the Old World porcupines of family Hystricidae, and the New World porcupines of family, Erethizont ...
s; * ''bristles'', which are long hairs usually used in visual signals, such as the mane of a lion; * ''velli'', often called "down fur", which insulates newborn mammals; and * '' wool'', which is long, soft, and often curly. Hair length is negligible in thermoregulation, as some tropical mammals, such as sloths, have the same fur length as some arctic mammals but with less insulation; and, conversely, other tropical mammals with short hair have the same insulating value as arctic mammals. The denseness of fur can increase an animal's insulation value, and arctic mammals especially have dense fur; for example, the musk ox has guard hairs measuring as well as a dense underfur, which forms an airtight coat, allowing them to survive in temperatures of . Some desert mammals, such as camels, use dense fur to prevent solar heat from reaching their skin, allowing the animal to stay cool; a camel's fur may reach in the summer, but the skin stays at . Aquatic mammals, conversely, trap air in their fur to conserve heat by keeping the skin dry. Mammalian coats are colored for a variety of reasons, the major selective pressures including camouflage, sexual selection, communication, and physiological processes such as temperature regulation. Camouflage is a powerful influence in many mammals, as it helps to conceal individuals from predators or prey.
Aposematism Aposematism is the advertising by an animal to potential predators that it is not worth attacking or eating. This unprofitability may consist of any defences which make the prey difficult to kill and eat, such as toxicity, venom, foul taste or ...
, warning off possible predators, is the most likely explanation of the black-and-white pelage of many mammals which are able to defend themselves, such as in the foul-smelling
skunk Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or ginge ...
and the powerful and aggressive honey badger. In arctic and subarctic mammals such as the arctic fox (''Vulpes lagopus''), collared lemming (''Dicrostonyx groenlandicus''), stoat (''Mustela erminea''), and snowshoe hare (''Lepus americanus''), seasonal color change between brown in summer and white in winter is driven largely by camouflage. Differences in female and male coat color may indicate nutrition and hormone levels, important in mate selection. Some arboreal mammals, notably primates and marsupials, have shades of violet, green, or blue skin on parts of their bodies, indicating some distinct advantage in their largely arboreal habitat due to convergent evolution. The green coloration of sloths, however, is the result of a
symbiotic Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
relationship with algae. Coat color is sometimes sexually dimorphic, as in many primate species. Coat color may influence the ability to retain heat, depending on how much light is reflected. Mammals with darker colored coats can absorb more heat from solar radiation and stay warmer; some smaller mammals, such as voles, have darker fur in the winter. The white, pigmentless fur of arctic mammals, such as the polar bear, may reflect more solar radiation directly onto the skin. The term ''pelage''first known use in English (French, from Middle French, from for 'hair', from Old French , from Latin )is sometimes used to refer to an animal's complete
coat A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
. The term fur is also used to refer to animal pelts which have been processed into leather with their hair still attached. The words ''fur'' or ''furry'' are also used, more casually, to refer to hair-like growths or formations, particularly when the subject being referred to exhibits a dense coat of fine, soft "hairs". If layered, rather than grown as a single Coat (animal), coat, it may consist of short down hairs, long wikt:guard hair, guard hairs, and in some cases, medium
awn hair Awn hairs are the intermediate hairs in a mammal's coat. They are shorter than the guard hairs and longer than the down hairs. They help with insulation and protect the down hairs underneath. Most of the visible coat is made of this kind of hair. ...
s. Mammals with reduced amounts of fur are often called "naked", as with the naked mole-rat, or "hairless", as with hairless dogs. An animal with commercially valuable fur is known within the Fur trade, fur industry as a List of types of fur, furbearer. The use of fur clothing, fur as clothing or decoration is controversial; animal welfare advocates object to the trapping and killing of wildlife, and to the confinement and killing of animals on Fur farming, fur farms.


Composition

The modern mammalian fur arrangement is known to have occurred as far back as Docodonta, docodonts, haramiyidans and Eutriconodonta, eutriconodonts, with specimens of ''Castorocauda'', ''Megaconus'' and ''Spinolestes'' preserving compound follicles with both guard hair and underfur. Fur may consist of three layers, each with a different type of hair.


Down hair

Down hair (also known as underfur, undercoat, underhair or ground hair) is the bottomor innerlayer, composed of wavy or curly hairs with no straight portions or sharp points. Down hairs, which are also flat, tend to be the shortest and most numerous in the coat. Thermoregulation is the principal function of the down hair, which insulates a layer of dry air next to the skin.


Awn hair

The
awn hair Awn hairs are the intermediate hairs in a mammal's coat. They are shorter than the guard hairs and longer than the down hairs. They help with insulation and protect the down hairs underneath. Most of the visible coat is made of this kind of hair. ...
can be thought of as a hybrid, bridging the gap between the distinctly different characteristics of down and guard hairs. Awn hairs begin their growth much like guard hairs, but less than half way to their full length, awn hairs start to grow thin and wavy like down hair. The Anatomical terms of location#Proximal and distal, proximal part of the awn hair assists in thermoregulation (like the down hair), whereas the Anatomical terms of location#Proximal and distal, distal part can shed water (like the guard hair). The awn hair's thin basal portion does not allow the amount of Goose bumps, piloerection that the stiffer guard hairs are capable of. Mammals with well developed down and guard hairs also usually have large numbers of awn hairs, which may even sometimes be the bulk of the visible coat.


Guard hair

Guard hair (overhair) is the top—or outer—layer of the coat. Guard hairs are longer, generally coarser, and have nearly straight shafts that protrude through the layer of softer down hair. The Anatomical terms of location, distal end of the guard hair is the visible layer of most mammal coats. This layer has the most marked pigmentation and Gloss (optics), gloss, manifesting as coat markings that are adapted for camouflage or display. Guard hair repels water and blocks sunlight, protecting the undercoat and skin in wet or aquatic habitats, and from the sun's ultraviolet radiation. Guard hairs can also reduce the severity of Wound#Open, cuts or scratches to the skin. Many mammals, such as the domestic dog and cat, have a pilomotor reflex that raises their guard hairs as part of a threat display when agitated.


Mammals with reduced fur

Hair is one of the defining characteristics of mammals; however, several species or breeds have considerably reduced amounts of fur. These are often called "naked" or "hairless".


Natural selection

Some mammals naturally have reduced amounts of fur. Some semiaquatic or aquatic animal, aquatic mammals such as cetaceans, pinnipeds and hippopotamuses have evolved hairlessness, presumably to reduce resistance through water. The naked mole-rat has evolved hairlessness, perhaps as an adaptation to their subterranean life-style. Two of the largest extant terrestrial mammals, the elephant and the rhinoceros, are largely hairless. The hairless bat is mostly hairless but does have short bristly hairs around its neck, on its front toes, and around the throat sac, along with fine hairs on the head and tail membrane. Most hairless animals cannot go in the sun for long periods of time, or stay in the cold for too long. Marsupial, Marsupials are born hairless and grow out fur later in development. Humans are the only primate species that have undergone significant hair loss. The hairlessness of humans compared to related species may be due to loss of functionality in the pseudogene KRTHAP1 (which helps produce keratin) Although the researchers dated the mutation to 240 000 ya, both the Altai Neandertal and Denisovan have the loss-of-function mutation, indicating it is much older. Mutations in the gene HR can lead to complete hair loss, though this is not typical in humans.


Artificial selection

At times, when a hairless domesticated animal is discovered, usually owing to a naturally occurring genetic mutation, humans may intentionally inbreed those hairless individuals and, after multiple generations, artificially create breeds that are hairless. There are several breeds of hairless cats, perhaps the most commonly known being the Sphynx (cat), Sphynx cat. Similarly, there are some breeds of hairless dogs. Other examples of artificially selected hairless animals include the Skinny pig, hairless guinea-pig, nude mouse, and the hairless rat.


Use in clothing

Fur has long served as a source of clothing for humans, including Neanderthals. Historically, it was worn for its insulating quality, with aesthetics becoming a factor over time. Pelts were worn in or out, depending on their characteristics and desired use. Today fur and trim used in garments may be dyed bright colors or to mimic exotic animal patterns, or shorn close like velvet. The term "a fur" may connote a coat, wrap, or shawl. The manufacturing of fur clothing involves obtaining animal pelts where the hair is left on the animal's processed skin. In contrast, making leather involves removing the hair from the hide or pelt and using only the skin. Fur is also used to make felt. A common felt is made from beaver fur and is used in bowler hats, top hats, and high-end cowboy hats.Chamber's journal, Published by Orr and Smith, 1952, p. 200, Original from the University of Michigan. List of types of fur, Common furbearers used include fox, rabbit, mink, leopard, beaver, Stoat, ermine, otter, sable, jaguar, pinniped, seal, coyote, chinchilla, raccoon, lemur, and Phalangeriformes, possum. File:Buckingham-palace-guard-11279634947G5ru.jpg, The iconic bearskins of the Queen's Guard at Buckingham Palace are made from the fur of American black bears. File:Carl Eielson.jpg, A fur seal, seal fur coat worn by Carl Ben Eielson (1897–1929), United States Air Force, USAF pilot and Arctic explorer File:Pelz-Verkaufsstand in Tallinn, Estland.jpg, A fur store in Tallinn, Estonia, in 2019


See also

*Angora wool *Animal coloration *Cat coat genetics *Coat (animal) *Plumage *Rabbit hair *Tanning (leather)


References


External links

{{Authority control Fur, Mammal anatomy Non-timber forest products