Description
Goatfish are characterized by two chin barbels (or goatee), which contain chemosensory organs and are used to probe the sand or holes in the reef for food. Their bodies are deep and elongated, with forked tails and widely separated dorsal fins. The first dorsal fin has 6-8 spines; the second dorsal has one spine and 8-9 soft rays, shorter than anal fin. Spines in anal fin 1 or 2, with 5-8 soft rays. They have 24 vertebrae. Many goatfish are brightly colored. The largest species, the dash-and-dot goatfish (''Parupeneus barberinus''), grows to in length; most species are less than half this size. Within the family are six Genus, genera and about 86 species.Genera
These genera are classified as belonging to the Mullidae: * ''Mulloidichthys'' Gilbert Percy Whitley, Whitley, 1929 * ''Mullus'' Carolus Linnaeus, Linnaeus, 1758 * ''Parupeneus'' Pieter Bleeker, Bleeker, 1863 * ''Pseudupeneus'' Bleeker, 1862 * ''Upeneichthys'' Bleeker, 1853 * ''Upeneus'' Georges Cuvier, Cuvier, 1829Distribution and habitat
Goatfish are distributed worldwide in tropical, subtropical, and temperate waters, in a range of habitats. Most species are associated with the bottom of the littoral zone, littoral, but some species of ''Upeneus'' can be deep; for example, the goatfish ''Upeneus davidaromi'' can be found at depths of . Tropical goatfish live in association with coral reefs. Some species, such as the freckled goatfish (''Upeneus tragula''), enter estuary, estuaries and rivers, although not to any great extent.Ecology
Goatfish are benthic feeders, using a pair of long chemosensory barbel (anatomy), barbels (whiskers) protruding from their chins to feel through the sediments in search of prey. They feed on worms, crustaceans, molluscs and other small invertebrates. Other fish shadow the active goatfish, waiting patiently for any overlooked prey. For example, in Indonesia large schools of the goldsaddle goatfish (''Parupeneus cyclostomus'') and moray eels hunt together. This behavior is known as shadow feeding or cooperative hunting. By day, many goatfish will form large Shoaling and schooling, schools of inactive (nonfeeding) fish; these aggregates may contain both conspecifics and heterospecifics. For example, the yellowfin goatfish (''Mulloidichthys vanicolensis'') is often seen congregating with bluestripe snappers (''Lutjanus kasmira''). All goatfish have the ability to change their coloration depending on their current activity. One notable example, the diurnal goldsaddle goatfish (''Parupeneus cyclostomus'') can change from a lemon-yellow to a pale cream whilst feeding.Mimicry
Goatfish have the ability to rapidly change color, and many species adopt a pale coloration when resting on the sand to blend with the background and become less visible to predators. These changes in color are reversible phenotypic changes and happen within seconds, many times during the lifespan of an individual. Two species, the mimic goatfish (''Mulloidichthys mimicus'') and Ayliffe's goatfish (''Mulloidichthys ayliffe'') have evolved to mimic the blue-striped snapper (''Lutjanus kasmira''), with which they often form schools. These are slow, genetic changes that have occurred during their evolution over many generations.Reproduction and lifecycle
Goatfish are pelagic spawners; they release many buoyant eggs into the water, which become part of the plankton. The eggs float freely with the currents until hatching. The larvae drift in oceanic waters or in the outer shelf for a period of 4–8 weeks until they metamorphose and develop barbels. Soon thereafter, most species take on a bottom-feeding lifestyle, although other species remain in the open water as juveniles or feed on plankton. Juvenile goatfish often prefer soft bottoms, in seagrass beds to mangroves. They change habitat preference as they develop, coinciding with changes in feeding habits, social behavior, and the formation of association with other species. Most species reach reproductive maturity after 1-2 years.Economic importance
Goatfish species are an important fishery in many areas of the world and some species are economically important. In ancient Rome until the end of the second century BCE, two species of goatfish (''Mullus barbatus'' and ''Mullus surmuletus'') were highly sought-after and expensive, not as a delicacy, but for aesthetic pleasure, since the fish assume a variety of colors and shades also during death. Therefore, it was paramount to serve the fish live and let them die before the eyes of the guests.Andrews, Alfred C. (1949). "The Roman Craze for Surmullets". ''The Classical Weekly'' 42 (12). Miami. 186–88.Timeline
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References
* {{Taxonbar, from=Q470850 Mullidae, Percoidei Taxa named by Constantine Samuel Rafinesque