Asian Elephant
   HOME

TheInfoList



OR:

The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is a species of elephant distributed throughout the Indian subcontinent and Southeast Asia, from India in the west to Borneo in the east, and Nepal in the north to
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
in the south. Three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised—'' E. m. maximus'', ''E. m. indicus'' and '' E. m. sumatranus''. The Asian elephant is characterised by its long
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
with a single finger-like processing; large tusks in males; laterally folded large ears but smaller in contrast to African elephants; and wrinkled grey skin. The skin is smoother than African elephants and may be
depigment Depigmentation is the lightening of the skin or loss of pigment. Depigmentation of the skin can be caused by a number of local and systemic conditions. The pigment loss can be partial (injury to the skin) or complete (caused by vitiligo). It can b ...
ed on the trunk, ears or neck. Adult males average in weight, and females . It is one of only three living species of elephants or elephantids anywhere in the world, the others being the
African bush elephant The African bush elephant (''Loxodonta africana'') is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to and a body ...
and African forest elephant and the only living species of the genus '' Elephas''. It is the second largest species of elephant after the African bush elephant. It frequently inhabits grasslands,  tropical evergreen forests, semi-evergreen forests, moist  deciduous forests, dry deciduous forests and dry thorn forests. They are herbivorous, eating about of vegetation per day. Cows and calves form groups, while males remain solitary or form "bachelor groups" with other males. During the breeding season, males will temporarily join female groups to mate. Asian elephants have a large and well-developed
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, sp ...
of the brain, are highly intelligent and self-aware being able to display behaviors associated grief, learning, greeting etc. The Asian elephant is the largest living land animal in Asia. Since 1986, the Asian elephant has been listed as Endangered on the IUCN Red List, as the population has declined by at least 50 per cent over the last three elephant generations, which is about 60–75 years. It is primarily threatened by
loss of habitat Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
, habitat degradation,
fragmentation Fragmentation or fragmented may refer to: Computers * Fragmentation (computing), a phenomenon of computer storage * File system fragmentation, the tendency of a file system to lay out the contents of files non-continuously * Fragmented distributi ...
and poaching. Wild Asian elephants live to be about 60 years old. While female captive elephants are recorded to have lived beyond 60 years when kept in semi-natural surroundings, Asian elephants die at a much younger age in captivity; captive populations are declining due to a low birth and high death rate. The earliest indications of captive use of Asian elephants are engravings on seals of the Indus Valley civilisation dated to the 3rd millennium BC.


Taxonomy

Carl Linnaeus proposed the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Elephas maximus'' in 1758 for an elephant from
Ceylon Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
. ''Elephas indicus'' was proposed by
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
in 1798, who described an elephant from India.
Coenraad Jacob Temminck Coenraad Jacob Temminck (; 31 March 1778 – 30 January 1858) was a Dutch people, Dutch Aristocracy (class), aristocrat, Zoology, zoologist and museum director. Biography Coenraad Jacob Temminck was born on 31 March 1778 in Amsterdam in the Dut ...
named an elephant from Sumatra ''Elephas sumatranus'' in 1847. Frederick Nutter Chasen classified all three as
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of the Asian elephant in 1940. These three subspecies are currently recognised as
valid Validity or Valid may refer to: Science/mathematics/statistics: * Validity (logic), a property of a logical argument * Scientific: ** Internal validity, the validity of causal inferences within scientific studies, usually based on experiments ** ...
taxa. Results of phylogeographic and morphological analyses indicate that the Sri Lankan and Indian elephants are not distinct enough to warrant classification as separate subspecies. Three subspecies are recognised: *
Sri Lankan elephant The Sri Lankan elephant (''Elephas maximus maximus'') is native to Sri Lanka and one of three recognised subspecies of the Asian elephant. It is the type subspecies of the Asian elephant and was first described by Carl Linnaeus under the bi ...
(''E. maximus maximus'' ) * Indian elephant (''E. maximus indicus'' ) * Sumatran elephant (''E. maximus sumatranus'' ) Sri Lankan elephants are the largest subspecies. Their skin colour is darker than of ''E. m. indicus'' and of ''E. m. sumatranus'' with larger and more distinct patches of depigmentation on ears, face, trunk and belly. The skin color of the Indian elephant is generally grey and lighter than that of ''E. m. maximus'' but darker than that of ''E. m. sumatranus''. A potential fourth subspecies, the
Borneo elephant The Borneo elephant, also called the Bornean elephant or the Borneo pygmy elephant, is a subspecies of Asian elephant ''(Elephas maximus)'' that inhabits northeastern Borneo, in Indonesia and Malaysia. Its origin remains the subject of debate. ...
(''Elephas maximus borneensis''), occurs in Borneo's northeastern parts, primarily in Sabah ( Malaysia), and sometimes in
Kalimantan Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo. In 2019, ...
( Indonesia). It was proposed by Paules Deraniyagala in 1950, who described an elephant in an illustration published in the ''
National Geographic ''National Geographic'' (formerly the ''National Geographic Magazine'', sometimes branded as NAT GEO) is a popular American monthly magazine published by National Geographic Partners. Known for its photojournalism, it is one of the most widely ...
'' magazine, but not as a living elephant in accordance with the rules of the International Code of Zoological Nomenclature. These elephants living in northern Borneo are smaller than all the other subspecies, but had larger ears, a longer tail, and straight tusks. Results of genetic analysis indicate that their ancestors separated from the mainland population about 300,000 years ago. A study in 2003, using
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
analysis and microsatellite data, indicated that the Borneo elephant population is derived from stock that originated in the region of the
Sunda Islands The Sunda Islands ( id, Kepulauan Sunda) are a group of islands in the Malay Archipelago.Chisholm, Hugh, ed. (1911). "Sunda Islands" . ''Encyclopædia Britannica''. 26 (11th ed.). Cambridge University Press. They consist of the Greater Sunda ...
, and suggests that the Borneo population has been separated from the other elephant populations of southeast Asia since the Pleistocene. The following Asian elephants were proposed as
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
subspecies, but are now considered
synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
with the Indian elephant: *
Syrian elephant The Syrian elephant or Western Asiatic elephant (''Elephas maximus asurus'') was the westernmost population of the Asian elephant (''Elephas maximus''), which became extinct in ancient times. Skeletal remains of ''E. m. asurus'' have been record ...
(''E. m. asurus''), proposed by Deraniyagala, based on fossil remains and Bronze Age illustrations. * Chinese elephant (''E. m. rubridens''), also proposed by Deraniyagala, based on a Chinese bronze statuette. * Javan elephant (''E. m. sondaicus''), also proposed by Deraniyagala, based on an illustration of a carving on the Buddhist monument of Borobudur.


Evolution

The genus '' Elephas'', of which the Asian elephant is the only living member, is the closest relative of the extinct mammoths. The two groups are estimated to have split from each other around 7 million years ago. ''Elephas'' originated in
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
during the Pliocene and spread throughout Africa before expanding into the southern half of Asia. The earliest ''Elephas'' species, '' Elephas ekorensis'', is known from the Early Pliocene of East Africa, around 5-4.2 million years ago. The oldest remains of the genus in Asia are known from the Siwalik Hills in the Indian subcontinent, dating to the late Pliocene, around 3.6-3.2 million years ago, assigned to the species '' Elephas planifrons.'' The modern Asian elephant is suggested to have evolved from the species '' Elephas hysudricus'', which first appeared at the beginning of the
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
around 2.6 million years ago, and is primarily known from remains of Early- Middle Pleistocene age found on the Indian subcontinent.' Skeletal remains of ''E. m. asurus'' have been recorded from the Middle East: Iran, Iraq,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
, and Turkey from periods dating between at least 1800 BC and likely 700 BC.


Description

In general, the Asian elephant is smaller than the
African bush elephant The African bush elephant (''Loxodonta africana'') is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to and a body ...
and has the highest body point on the head. The back is convex or level. The ears are small with dorsal borders folded laterally. It has up to 20 pairs of ribs and 34
caudal vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
. The feet have five nail-like structures on each forefoot, and four on each hind foot. The forehead has two hemispherical bulges, unlike the flat front of the African elephants. Its long
trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...
or proboscis has only one fingerlike tip, in contrast to the African elephants which have two. Hence, the Asian species relies more on wrapping around a food item and squeezing it into its mouth, rather than grasping with the tip. Asian elephants have more muscle coordination and can perform more complex tasks.Cows usually lack tusks; if tusks—in that case, called "tushes"—are present, they are barely visible and only seen when the mouth is open. The enamel plates of the molars are greater in number and closer together in Asian elephants. Some bulls may also lack tusks; these individuals are called "makhnas", and are especially common among the Sri Lankan elephant population. A tusk from an tall elephant killed by Sir
Victor Brooke Sir Victor Alexander Brooke, 3rd Baronet (5 January 1843 – 23 November 1891), was an Anglo-Irish sportsman- naturalist and baronet. He was the father of Field Marshal The 1st Viscount Alanbrooke, and grandfather of The 1st Viscount Brookeboro ...
measured in length, and nearly in circumference, and weighed . This tusk's weight is, however, exceeded by the weight of a shorter tusk of about in length which weighed , and there have reportedly been tusks weighing over . Skin colour is usually grey, and may be masked by soil because of dusting and wallowing. Their wrinkled skin is movable and contains many nerve centres. It is smoother than that of African elephants and may be depigmented on the trunk, ears, or neck. The
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rele ...
and
dermis The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided i ...
of the body average thick; skin on the dorsum is thick providing protection against bites, bumps, and adverse weather. Its folds increase surface area for heat dissipation. They can tolerate cold better than excessive heat. Skin temperature varies from . Body temperature averages .


Size

On average, when fully-grown, bulls are about tall at the shoulder and in weight, while cows are smaller at about at the shoulder and in weight. Sexual dimorphism in body size is relatively less pronounced in Asian elephants than in African bush elephants; with bulls averaging 15% and 23% taller in the former and latter respectively. Length of body and head including trunk is with the tail being long. The largest bull elephant ever recorded was shot by the Maharajah of Susang in the Garo Hills of Assam, India, in 1924, it weighed an estimated , stood tall at the shoulder and was long from head to tail. There are reports of larger individuals as tall as .


Distribution and habitat

Asian elephants are distributed throughout the  Indian subcontinent and  Southeast Asia, from  India in the west, to  Borneo in the east, and Nepal in the north, to
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
 in the south. They inhabit grasslands, tropical evergreen forests, semi-evergreen forests, moist deciduous forests, dry deciduous forests and dry thorn forests, in addition to cultivated and secondary forests and scrublands. Over this range of habitat types elephants occur from sea level to over . In the eastern Himalaya in northeast India, they regularly move up above in summer at a few sites. In Bangladesh, some isolated populations survived in the south-east
Chittagong Hills The Chittagong Hill Tracts ( bn, পার্বত্য চট্টগ্রাম, Parbotto Chottogram), often shortened to simply the Hill Tracts and abbreviated to CHT, are group of districts within the Chittagong Division in southeast ...
in the early 1990s. In Malaysia's northern Johor and Terengganu National Park, two Asian elephants tracked using satellite tracking technology spent most of their time in secondary or "logged-over forest"; they travelled 75% of their time in an area less than away from a water source. In
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, the Asian elephant survives only in the prefectures of Xishuangbanna, Simao and Lincang of southern Yunnan. , the estimated population was around 300 individuals. As of 2017, the estimated wild population in India account for nearly three-fourths of the extant population, at 27,312 individuals. In 2019, the Asian elephant population in India increased to an estimated 27,000–29,000 individuals. , the global wild population was estimated at 48,323–51,680 individuals.


Ecology and behaviour

Asian elephants are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
. They are classified as megaherbivores and consume up to of plant matter per day. Around 50 to 75% of the day is devoted to eating. They are generalist feeders, and are both grazers and browsers. They are known to feed on at least 112 different plant species, most commonly of the order Malvales, as well as the
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...
, palm, sedge and
true grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
families. They browse more in the dry season with bark constituting a major part of their diet in the cool part of that season. They drink at least once a day and are never far from a permanent source of fresh water. They need 80–200 litres of water a day and use even more for bathing. At times, they scrape the soil for clay or minerals. Cows and calves move about together as groups, while bulls disperse from their mothers upon reaching adolescence. Bulls are solitary or form temporary "bachelor groups". Cow-calf units generally tend to be small, typically consisting of three adults (most likely related females) and their offspring. Larger groups of as many as 15 adult females have also been recorded. Seasonal aggregations of 17 individuals including calves and young adults have been observed in Sri Lanka's
Uda Walawe National Park Udawalawe National Park is a national park on the boundary of Sabaragamuwa and Uva Provinces in Sri Lanka. The park was created to provide a sanctuary for wild animals displaced by the construction of the Udawalawe Reservoir on the Walawe River ...
. Until recently, Asian elephants, like African elephants, were thought to be under the leadership of older adult females, or matriarchs. It is now recognized that cows form extensive and very fluid social networks, with varying degrees of associations between individuals. Social ties generally tend to be weaker than in African bush elephants. Unlike African elephants, which rarely use their forefeet for anything other than digging or scraping soil, Asian elephants are more agile at using their feet in conjunction with the trunk for manipulating objects. They can sometimes be known for their violent behavior. Asian elephants are recorded to make three basic sounds: growls, squeaks and snorts. Growls in their basic form are used for short distance communication. During mild arousal, growls resonate in the trunk and become rumbles while for long-distance communication, they escalate into roars. Low-frequency growls are infrasonic and made in many contexts. Squeaks come in two forms: chirpings and trumpets. Chirping consists of multiple short squeaks and signals conflict and nervousness. Trumpets are lengthened squeaks with increased loudness and are produced during extreme arousal. Snorts signal changes in activity and increase in loudness during mild or strong arousal. During the latter case, when an elephant bounces the tip of the trunk, it creates booms which serve as threat displays. Elephants can distinguish low-amplitude sounds. Rarely, tigers have been recorded attacking and killing calves, especially if the calves become separated from their mothers, stranded from their herd, or orphaned. Adults are largely invulnerable to natural predation. There is a singular anecdotal case of a mother Asian elephant allegedly being killed alongside her calf; however, this account is contestable. In 2011 and 2014, two instances were recorded of tigers successfully killing adult elephants; one by a single tiger in Jim Corbett National Park on a 20-year-old young adult elephant cow, and another on a 28-year-old sick adult bull in Kaziranga National Park further east, which was taken down and eaten by several tigers hunting cooperatively. Elephants appear to distinguish between the growls of larger predators like tigers and smaller predators like
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
s; they react to leopards less fearfully and more aggressively.


Reproduction

Reproduction in Asian elephants can be attributed to the production and perception of signaling compounds called pheromones. These signals are transmitted through various bodily fluids. They are commonly released in urine but in males they are also found in special secretions from the temporal glands. Once integrated and perceived, these signals provide the receiver with information about the reproductive status of the sender. If both parties are ready to breed, reproductive ritualic behavior occurs and the process of sexual reproduction proceeds. Bulls will fight one another to get access to oestrus cows. Strong fights over access to females are extremely rare. Bulls reach sexual maturity around the age of 12–15. Between the ages of 10 and 20 years, bulls undergo an annual phenomenon known as " musth". This is a period where the testosterone level is up to 100 times greater than non-musth periods, and they become aggressive. Secretions containing pheromones occur during this period, from the paired temporal glands located on the head between the lateral edge of the eye and the base of the ear. The aggressive behaviors observed during musth can be attributed to varying amounts of frontalin (1,5-dimethyl-6,8-dioxabicyclo .2.1ctane) throughout the maturation process of bulls. Frontalin is a pheromone that was first isolated in bark beetles but can also be produced in the bulls of both Asian and African Elephants. The compound can be excreted through urine as well as through the temporal glands of the bull, allowing signaling to occur. During musth, increased concentrations of frontalin in the bull's urine communicate the reproductive status of the bull to female elephants. Similar to other mammals, hormone secretion in female elephants is regulated by an estrous cycle. This cycle is regulated by surges in Luteinizing hormone that are observed three weeks from each other. This type of estrous cycle has also been observed in African Elephants but is not known to affect other mammals. The first surge in Luteinizing hormone is not followed by the release of an egg from the ovaries. However, some female elephants still exhibit the expected mating protocols during this surge. Female elephants give ovulatory cues by utilizing sex pheromones. A principal component thereof, (Z)-7-dodecen-1-yl acetate, has also been found to be a sex pheromone in numerous species of insects. In both insects and elephants, this chemical compound is used as an attractant to assist the mating process. In elephants, the chemical is secreted through urination and this aids in the attraction of bulls to mate. Once detected, the chemical stimulates the vomeronasal organ of the bull, thus providing information on the maturity of the female. Reproductive signaling exchange between male and female elephants are transmitted through olfactory cues in bodily fluids. In males, the increase in frontalin during musth heightens their sensitivity to the (Z)-7-dodecen-1-yl acetate produced by female elephants. Once perceived by receptors in the trunk, a sequence of ritualistic behaviors follow. The responses in males vary based on both the stage of development and the temperament of the elephant. This process of receiving and processing signals through the trunk is referred to as flehmen. The difference in body movements give cues to gauge if the male is interested in breeding with the female that produced the secretion. A bull that is ready to breed will move closer to the urine and in some cases an erection response is elicited. A bull that is not ready to breed will be timid and try to dissociate themselves from the signal. In addition to reproductive communication, chemosensory signaling is used to facilitate same-sex interactions. When less developed males detect pheromones from a male in musth, they often retreat to avoid coming in contact with aggressive behaviors. Female elephants have also been seen to communicate with each other through pheromone in urine. The purpose of this type of intersex communication is still being investigated. However, there are clear differences in signaling strength and receiver response throughout different stages of the estrous cycle. The
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once it ...
is 18–22 months, and the cow gives birth to one calf, only occasionally twins. The calf is fully developed by the 19th month, but stays in the womb to grow so that it can reach its mother to feed. At birth, the calf weighs about , and is
suckled Breastfeeding, or nursing, is the process by which human breast milk is fed to a child. Breast milk may be from the breast, or may be expressed by hand or pumped and fed to the infant. The World Health Organization (WHO) recommends that brea ...
for up to three years. Once a female gives birth, she usually does not breed again until the first calf is weaned, resulting in a four to five-year birth interval. During this period, mother to calf communication primarily takes place through temporal means. However, male calves have been known to develop sex pheromone-producing organs at a young age. Early maturity of the vomeronasal organ allows immature elephants to produce and receive pheromones. It is unlikely that the integration of these pheromones will result in a flehmen response in a calf. Females stay on with the herd, but mature males are chased away. Female Asian elephants sexually mature around the age of 10~15 and keep growing until 30, while males fully mature at more than the age of 25, and constantly grow throughout their life. Average elephant life expectancy is 60 years in the wild and 80 in captivity, although this has been exaggerated in the past. Generation length of the Asian elephant is 22 years.


Intelligence

Asian elephants have a very large and highly developed
neocortex The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, sp ...
, a trait also shared by humans,
apes Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...
and certain dolphin species. They have a greater volume of cerebral cortex available for
cognitive Cognition refers to "the mental action or process of acquiring knowledge and understanding through thought, experience, and the senses". It encompasses all aspects of intellectual functions and processes such as: perception, attention, thought, ...
processing than all other existing land animals. Results of studies indicate that Asian elephants have cognitive abilities for tool use and tool-making similar to great apes. They exhibit a wide variety of behaviours, including those associated with
grief Grief is the response to loss, particularly to the loss of someone or some living thing that has died, to which a bond or affection was formed. Although conventionally focused on the emotional response to loss, grief also has physical, cogni ...
, learning, allomothering,
mimicry In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. Often, mimicry f ...
, play,
altruism Altruism is the principle and moral practice of concern for the welfare and/or happiness of other human beings or animals, resulting in a quality of life both material and spiritual. It is a traditional virtue in many cultures and a core as ...
, use of
tools A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ba ...
,
compassion Compassion motivates people to go out of their way to relieve the physical, mental or emotional pains of others and themselves. Compassion is often regarded as being sensitive to the emotional aspects of the suffering of others. When based on n ...
,
cooperation Cooperation (written as co-operation in British English) is the process of groups of organisms working or acting together for common, mutual, or some underlying benefit, as opposed to working in competition for selfish benefit. Many animal a ...
,
self-awareness In philosophy of self, self-awareness is the experience of one's own personality or individuality. It is not to be confused with consciousness in the sense of qualia. While consciousness is being aware of one's environment and body and lifesty ...
, memory, and language. Elephants reportedly head to safer ground during natural disasters like tsunamis and earthquakes, but data from two satellite-collared Sri Lankan elephants indicate this may be untrue. Several students of elephant cognition and neuroanatomy are convinced that Asian elephants are highly intelligent and self-aware. Others contest this view.


Threats

The pre-eminent threats to the Asian elephant today are the loss, degradation and fragmentation of its habitat, which leads to increasing conflicts between humans and elephants. Asian elephants are poached for ivory and a variety of other products including
meat Meat is animal flesh that is eaten as food. Humans have hunted, farmed, and scavenged animals for meat since prehistoric times. The establishment of settlements in the Neolithic Revolution allowed the domestication of animals such as chic ...
and leather. The demand for elephant skin has risen due to it being an increasingly-common ingredient in traditional Chinese medicine.


Human–elephant conflict

In some parts of Asia, people and elephants have co-existed for thousands of years. In other areas, people and elephants come into conflict, resulting in violence, and ultimately, the displacement of elephants. The main causes of human-elephant conflict includes the growing human population, large-scale development projects and poor top-down governance. Proximate causes includes habitat loss due to deforestation, disruption of elephant migratory routes, expansion of agriculture and illegal encroachment into protected areas. Destruction of forests through
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidding, on-site processing, and loading of trees or logs onto trucks or skeleton cars. Logging is the beginning of a supply chain ...
, encroachment, slash-and-burn, shifting cultivation, and monoculture tree plantations are major threats to the survival of elephants. Human–elephant conflicts occur when elephants raid crops of shifting cultivators in fields, which are scattered over a large area interspersed with forests. Depredation in human settlements is another major area of human–elephant conflict occurring in small forest pockets, encroachments into elephant habitat, and on elephant migration routes. However, studies in Sri Lanka indicate that traditional slash-and-burn agriculture may create optimal habitat for elephants by creating a mosaic of successional-stage vegetation. Populations inhabiting small habitat fragments are much more liable to come into conflict with humans. Development such as border fencing along the India–Bangladesh border has become a major impediment to the free movement of elephants. In Assam, more than 1,150 humans and 370 elephants died as a result of human-elephant conflict between 1980 and 2003. In a 2010 study, it was estimated that in India alone, over 400 people were killed by elephants each year, and 0.8 to 1 million hectares were damaged, affecting at least 500,000 families across the country.Rangarajan, M., Desai, A., Sukumar, R., Easa, P. S., Menon, V., Vincent, S., Ganguly, S., Talukdar, B. K., Singh, B., Mudappa, D., Chowdhary, S., Prasad, A. N. (2010)
''Gajah: Securing the future for elephants in India''
. Report of the Elephant Task Force. Ministry of Environment and Forests, New Delhi.
Moreover, elephants are known to destroy crops worth up to US$2–3 million annually. This has major impacts on the welfare and livelihoods of local communities, as well as the future conservation of this species. In countries like Bangladesh and
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, the Asian elephant is one of the most feared wild animals, even though they are less deadly than other local animals such as venomous snakes (which were estimated to claim more than 30 times more lives in Sri Lanka than elephants). As a whole, Asian elephants display highly sophisticated and sometimes unpredictable behaviour. Most untamed elephants try to avoid humans, but if they are caught off guard by any perceived physical threat, including humans, they will likely charge. This is especially true of males in musth and of females with young. Gunfire and other similar methods of deterring, which are known to be effective against many kinds of wild animals including tigers, may or may not work with elephants, and can even worsen the situation. Elephants that have been abused by humans in the past often become "rogue elephants", which regularly attack people with no provocation.


Poaching


For ivory

The demand for ivory during the 1970s and 1980s, particularly in East Asia, led to rampant poaching and the serious decline of elephants in both Africa and Asia. In Thailand, the illegal trade in live elephants and ivory still flourishes. Although the amount of ivory being openly sold has decreased substantially since 2001, Thailand still has one of the largest and most active black markets for ivory seen anywhere in the world. Tusks from Thai-poached elephants also enter the market; between 1992 and 1997 at least 24 male elephants were killed for their tusks.Stiles, D. (2009)
''The elephant and ivory trade in Thailand.''
TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia.
Up to the early 1990s, Vietnamese ivory craftsmen used exclusively Asian elephant ivory from Vietnam and neighbouring Lao and Cambodia. Before 1990, there were few tourists and the low demand for worked ivory could be supplied by domestic elephants. Economic liberalisation and an increase in tourism raised both local and visitors' demands for worked ivory, which resulted in heavy poaching.


For skin

The skin of the Asian elephant is used as an ingredient in Chinese medicine as well as in the manufacture of ornamental beads. The practice has been aided by China's
State Forestry Administration The State Forestry and Grassland Administration () is an administration of the People's Republic of China, in charge of the national forestry affairs. It was founded in 1949 as Ministry of Forestry and Farming. On November 5, 1951, it changed to ...
(SFA), which has issued licences for the manufacture and sale of pharmaceutical products containing elephant skin, thereby making trading legal. In 2010, four skinned elephants were found in a forest in Myanmar; 26 elephants were killed by poachers in 2013 and 61 in 2016. According to the NGO Elephant Family, Myanmar is the main source of elephant skin, where a poaching crisis has developed rapidly since 2010.


Disease

The
Elephant endotheliotropic herpesvirus Elephant endotheliotropic herpesviruses (EEHV) or ''Elephantid betaherpesvirus 1'' (ElHV-1) is a type of herpesvirus, which can cause a highly fatal hemorrhagic disease when transmitted to young Asian elephants. In African elephants, related form ...
(EEHV) is a member of the ''
Proboscivirus ''Proboscivirus'' is a genus of viruses in the order ''Herpesvirales'', in the family ''Herpesviridae'', in the subfamily ''Betaherpesvirinae''. Elephants serve as natural hosts. EEHV1 is apathogenic for African elephants but causes fatal haemo ...
'' genus, a novel clade most closely related to the mammalian
betaherpesviruses ''Betaherpesvirinae'' is a subfamily of viruses in the order '' Herpesvirales'' and in the family '' Herpesviridae''. Mammals serve as natural hosts. There are 26 species in this subfamily, divided among 5 genera. Diseases associated with this s ...
. As of 2011, it is responsible for as many as 70 deaths of both zoo and wild Asian elephants worldwide, especially in young calves. In particular, several incidents of calves dying from
elephant endotheliotropic herpesvirus Elephant endotheliotropic herpesviruses (EEHV) or ''Elephantid betaherpesvirus 1'' (ElHV-1) is a type of herpesvirus, which can cause a highly fatal hemorrhagic disease when transmitted to young Asian elephants. In African elephants, related form ...
have been recorded in Myanmar. The elephant schistosome is a parasitic trematode that uses the Asian elephant as a definitive host. Two other hosts may be the Indian elephant and the greater one-horned rhinoceros.


Conservation

The Asian elephant is listed on CITES Appendix I. It is a quintessential flagship species, deployed to catalyze a range of conservation goals, including habitat conservation at landscape scales, generating public awareness on conservation issues, and mobilisation as a popular cultural icon both in India and the West. A key aspect of conservation is connectivity of the preferred movement routes of Asian elephants through areas with high vegetation cover and low human population density. The
World Elephant Day World Elephant Day is an international annual event on August 12, dedicated to the preservation and protection of the world's elephants. Conceived in 2011 by Canadian filmmakers Patricia Sims and Michael Clark of Canazwest Pictures, and Sivaporn D ...
is celebrated annually on 12 August since 2012. Events are organized to divulge information and to engage people about the problems that the Asian elephant is facing. August has been established as the Asian Elephant Awareness Month by zoos and conservation partners in the United States.
Karnataka state Karnataka (; ISO: , , also known as Karunāḍu) is a state in the southwestern region of India. It was formed on 1 November 1956, with the passage of the States Reorganisation Act. Originally known as Mysore State , it was renamed ''Karnat ...
in India, hosts the most Asian elephants of any known area and comprises 20% of the total population in the country. The distribution of elephants in the state according to one estimate is roughly . In a 2013 study, an estimated 10, 000 elephants inhabited the Western Ghats, and were primarily threatened by poaching and habitat fragmentation. An increase in conflict with humans was also sited as a likely issue. Conservation plans aim to establish wildlife corridors, stop poaching of bulls, and protect or manage land area. Project Elephant was initiated in 1992 as a Centrally Sponsored Scheme (CSS) by the Ministry of Environment, Forest and Climate Change of the Government of India. The project was initiated to protect the Indian elephant and its habitats and to establish dedicated elephant reserves for sustaining elephant populations. The distribution of elephants in Sri Lanka is only two-fifths of what it was in the late 19th and early 20th centuries. Thus, interactions with humans occur much more frequently. During a 2003 survey, the local people expressed some form of disapproval towards the conservation of Asian elephants as farmers viewed them as pests, however, most of the participants were supportive of the idea. In China, Asian elephants are under first-level protection. Yunnan province has 11 national and regional nature reserves. In total, the covered
protected area in China This is a list of the nationally-designated protected areas of China. There are many forms of protected areas in China. Based on their relative importance, each type of protected area can be further graded into two to three levels (national, pro ...
is about . In 2020, the population of Asian elephants in Yunnan was estimated at around 300 individuals. As conflicts between humans and wild elephants have emerged around protected areas in the last years, the prefecture of Xishuangbanna built food bases and planted bananas and bamboo to create a better habitat. In Thailand, Salakpra Wildlife Sanctuary and
Tham Than Lot National Park Chaloem Rattanakosin National Park ( th, อุทยานแห่งชาติเฉลิมรัตนโกสินทร์), also known as Tham Than Lot National Park (อุทยานแห่งชาติถ้ำธารลอ ...
are protected areas hosting around 250–300 elephants, according to figures from . In recent years the National Park has faced issues due to encroachment and over-exploitation. In India, the
National Board of Wildlife National may refer to: Common uses * Nation or country ** Nationality – a ''national'' is a person who is subject to a nation, regardless of whether the person has full rights as a citizen Places in the United States * National, Maryland, ce ...
recommended to allow coal mining in
Dehing Patkai National Park Dehing Patkai National Park is located in the Dibrugarh district, Dibrugarh and Tinsukia districts of Assam and covers an area of rainforest. It was declared a wildlife sanctuary on 13 June 2004. On 13 December 2020 Government of Assam upgraded ...
in April 2020. The decision raised concerns between students and environmental activists who launched an online campaign to stop the project.


In captivity

About half of the global zoo elephant population is kept in European zoos, where they have less than half (18.9 years) the median life span of conspecifics (41.6 years) in protected populations in range countries. This discrepancy is clearest in Asian elephants: infant mortality is more than two to three times that seen in Burmese timber camps, and adult survivorship in zoos has not improved significantly in recent years. One risk factor for Asian zoo elephants is being moved between institutions, with early removal from the mother tending to have additional adverse effects. Another risk factor is being born into a zoo rather than being imported from the wild, with poor adult survivorship in zoo-born Asians apparently being conferred prenatally or in early infancy. Likely causes for compromised survivorship is stress and/or obesity. Foot problems are commonly observed in captive elephants. These are related to lack of exercise, long hours standing on hard substrates, and contamination resulting from standing in their dung. Many of these problems are treatable. However, mistreatment may lead to serious disability or death. Demographic analysis of captive Asian elephants in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
indicates that the population is not self-sustaining. First year mortality is nearly 30 per cent, and fecundity is extremely low throughout the prime reproductive years. Data from North American and European regional studbooks from 1962 to 2006 were analysed for deviations in the birth and juvenile death sex ratios. Of 349 captive calves born, 142 died prematurely. They died within one month of birth, major causes being
stillbirth Stillbirth is typically defined as fetal death at or after 20 or 28 weeks of pregnancy, depending on the source. It results in a baby born without signs of life. A stillbirth can result in the feeling of guilt or grief in the mother. The term ...
and
infanticide Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of reso ...
by either the calf's mother or by one of the exhibition mates. The sex ratio of stillbirths in Europe was found to have a tendency for excess of males.


Handling methods

Young elephants are captured from the wild and illegally imported to Thailand from
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
for use in the tourism industry; calves are used mainly in amusement parks and are trained to perform various stunts for tourists. The calves are often subjected to a 'breaking in' process, which may involve being tied up, confined, starved, beaten and tortured; as a result, two-thirds may perish. Handlers use a technique known as the
training crush Elephant crushing, or a training crush, is a method by which wild elephants can be tamed for domestication, using restriction in a cage, sometimes with the use of corporal punishment or negative reinforcement. This practice is condemned by a va ...
, in which "handlers use sleep-deprivation, hunger, and thirst to "break" the elephants' spirit and make them submissive to their owners"; moreover, handlers drive nails into the elephants' ears and feet.


In culture

The Asian elephant is the national animal of Thailand and Laos. It has also been declared as the national heritage animal of India. Bones of Asian elephants excavated at Mohenjo-daro in the
Indus Valley The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmir, ...
indicate that they were tamed in the Indus Valley Civilisation and used for work. Decorated elephants are also depicted on seals and were modelled in clay. The Asian elephant became a siege engine, a mount in war, a
status symbol A status symbol is a visible, external symbol of one's social position, an indicator of economic or social status. Many luxury goods are often considered status symbols. ''Status symbol'' is also a sociological term – as part of social and soci ...
, a beast of burden, and an elevated platform for hunting during historical times in South Asia. Asian elephants have been captured from the wild and tamed for use by humans. Their ability to work under instruction makes them particularly useful for carrying heavy objects. They have been used particularly for timber-carrying in jungle areas. Other than their work use, they have been used in war, in ceremonies, and for carriage. It is reported that war elephants are still used by the Kachin Independence Army (KIA) in
Kachin State Kachin State ( my, ကချင်ပြည်နယ်; Kachin: ), also known by the endonym Kachinland, is the northernmost state of Myanmar. It is bordered by China to the north and east (Tibet and Yunnan, specifically and respectively); Sh ...
in northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
against Myanmar's military. The KIA use about four dozen elephants to carry supplies. The Asian elephant plays an important part in the culture of the subcontinent and beyond, being featured prominently in the ''
Panchatantra The ''Panchatantra'' (IAST: Pañcatantra, ISO: Pañcatantra, sa, पञ्चतन्त्र, "Five Treatises") is an ancient Indian collection of interrelated animal fables in Sanskrit verse and prose, arranged within a frame story.
'' fables and the Buddhist '' Jataka'' tales. They play a major role in Hinduism: the god Ganesha's head is that of an elephant, and the "blessings" of a temple elephant are highly valued. Elephants are frequently used in processions where the animals are adorned with festive outfits. The Asian elephant is depicted in several Indian manuscripts and treatises with notable amongst these including ''
Matanga Lila Matanga Lila is a treatise in Sanskrit dealing with the life and behaviour of elephants. The title has been translated as Elephant-sport. It is a treatise in 263 stanzas divided into twelve chapters of varying length. In the treatise, the author ...
'' (elephant sport) of Nilakantha. The manuscript ''
Hastividyarnava The Hastividyārnava, written by Sukumar Barkaith, is one of the best known illustrated manuscripts of Assam. Commissioned under the patronage of King Siva Singha (1714-1744 C.E.) and his queen consort Ambika, it deals with the management and car ...
'' is from Assam in northeast India. In the Burmese, Thai and Sinhalese animal and planetary zodiac, the Asian elephant, both tusked and tuskless, are the fourth and fifth animal zodiacs of the
Burmese Burmese may refer to: * Something of, from, or related to Myanmar, a country in Southeast Asia * Burmese people * Burmese language * Burmese alphabet * Burmese cuisine * Burmese culture Animals * Burmese cat * Burmese chicken * Burmese (hor ...
, the fourth animal zodiac of the
Thai Thai or THAI may refer to: * Of or from Thailand, a country in Southeast Asia ** Thai people, the dominant ethnic group of Thailand ** Thai language, a Tai-Kadai language spoken mainly in and around Thailand *** Thai script *** Thai (Unicode block ...
, and the second animal zodiac of the Sinhalese people of Sri Lanka. Similarly, the elephant is the twelfth animal zodiac in the Dai animal zodiac of the
Dai people The Dai people ( Burmese: ရှမ်းလူမျိုး; khb, ᨴᩱ/ᨴᩱ᩠ᨿ; lo, ໄຕ; th, ไท; shn, တႆး, ; , ; ) refers to several Tai-speaking ethnic groups living in the Xishuangbanna Dai Autonomous Prefecture and t ...
in southern China.


See also

* Elephants in Thailand *
Endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
*
Ivory trade The ivory trade is the commercial, often illegal trade in the ivory tusks of the hippopotamus, walrus, narwhal, mammoth, and most commonly, African and Asian elephants. Ivory has been traded for hundreds of years by people in Africa and Asia, ...
* Khedda * List of individual elephants * Mela shikar * War elephant * White elephant


References


Bibliography

*


Further reading

* *


External links


Save Elephant Foundation

International Elephant Foundation

ElefantAsia: ''Protecting the Asian elephant''

Asian Elephants at the Zoological Gardens of the World

Elephant Information Repository

WWF—Asian elephant species profile

National Zoo Facts on Asian Elephant and a Webcam of the Asian Elephant exhibit

Environmental Investigation Agency: ''Illegal Wildlife Trade'' : ''Elephants''
{{Authority control Elephants EDGE species Endangered animals Endangered biota of Asia Extant Pliocene first appearances Fauna of South Asia Fauna of Southeast Asia Herbivorous mammals Mammals described in 1758 Mammals of Asia National symbols of India National symbols of Laos National symbols of Thailand Pliocene mammals of Asia Quaternary animals of Asia Species endangered by habitat fragmentation Species endangered by habitat loss Taxa named by Carl Linnaeus