Zygadenia
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''Zygadenia'' is an extinct genus of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s in the family
Ommatidae The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. How ...
.


Triassic Species

After Kirejtshuk, 2020. Kirejtshuk notes that the two Lower Jurassic Australian species ''Z. westraliensis'' and ''Z. martinae'' possibly merit placement in their own separate genus.


Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
(242 Ma to ~237 Ma)

; Tongchuan Formation, China * ''Z. shiluoensis''


Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
(237 Ma to 227 Ma)

; Koldzat Formation, Kazakhstan * ''Z. oxypygus'' ;
Madygen Formation The Madygen Formation ( Russian: Madygen Svita) is a Late Triassic (Carnian) geologic formation and Lagerstätte in the Batken and Osh Regions of western Kyrgyzstan, with minor outcrops in neighboring Tajikistan and Uzbekistan. The conglomerates, ...
, Kyrgyzstan * ''Z. laticella'' * ''Z. rostratus'' * ''Z. tenuis''


Jurassic species


Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
(201.3 Ma to 199.3 Ma)

; Dzhil Formation, Kyrgyzstan * ''Z. cellulosa'' * ''Z. issykkulensis'' * ''Z. kirghizicus'' * ''Z. latus'' * ''Z. sogutensis'' ; Guanyintan Formation, China * ''Z. homorus''


Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
(182.7 Ma to 174.1 Ma)

;
Cattamarra Coal Measures The Cattamarra Coal Measures is an Early Jurassic geological unit in Western Australia. Description They are part of the Perth Basin, and are a sequence of non-marine, probably fluvial sandstones, shales and silts including bituminous coal, and ...
, Australia * ''Z. martinae'' * ''Z. westraliensis'' ; Guanyintan Formation, China * ''Z. lini'' ; Sulyukta Formation, Kyrgyzstan * ''Z. crassus''


Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratig ...
(170.3 Ma to 168.3 Ma)

; Bakhar Formation, Mongolia * ''Z. brachycephalus'' * ''Z. exiguus'' * ''Z. longicollis''


Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
(166.1 Ma to 163.5 Ma)

;
Haifanggou Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...
, China * ''Z. dischides'' * ''Z. jurassica'' * ''Z. pingi'' ; Tyumen Formation, Russian * ''Z. patulus''


Oxfordian (163.5 Ma to 157.3 Ma)

;
Karabastau Formation The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...
, Kazakhstan * ''Z. foersteri'' * ''Z. lapidarius'' * ''Z. nigromonticola'' * ''Z. picturatus'' * ''Z. pulcher''


Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
(152.1 Ma To 145.0 Ma)

; Shar Teeg, Mongolia * ''Z. brachycephalus'' * ''Z. exiguus'' * ''Z. giebeli'' * ''Z. handlirschi'' ;
Solnhofen Solnhofen is a municipality in the district of Weißenburg-Gunzenhausen in the region of Middle Franconia in the ' of Bavaria in Germany. It is in the Altmühl valley. The local area is famous in geology and palaeontology for Solnhofen limest ...
, Germany * ''Z. reticulatus'' ) * ''Z. tripartitus''


Cretaceous species


Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...
(145.0 Ma To 139.8 Ma)

;
Lulworth Formation The Lulworth Formation is a Formation (geology), geologic formation in England. It dates from the late Tithonian to the mid Berriasian. It is a subunit of the Purbeck Group. In Dorset, it consists of three members, which are in ascending order, t ...
, United Kingdom * ''Z. angliae'' * ''Z. tuberculata''


Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.9 ± 2 Ma and 129.4 ± 1.5 Ma (million years ago). The Hauterivian is preceded by the Va ...
(132.9 Ma to 129.4 Ma)

;
Dabeigou Formation The Dabeigou Formation is a paleontology, palaeontological formation located in Hebei, China.Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ro ...
, United Kingdom * ''Z. angliae'' * ''Z. floodpagei'' * ''Z. simpsoni''


Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
(129.4 Ma to 125.0 Ma)

; Jianshangou Formation, China * ''Z. laetus'' ;
La Huérguina Formation The La Huérguina Formation (also known as the Calizas de La Huérguina Formation, La Huérguina Limestone Formation or as the Una Formation) is a geological formation in Spain whose strata date back to the Barremian stage of the Early Cretaceous ...
, Las Hoyas, Spain * ''Z. diazromerali'' * ''Z. longicoxa'' * ''Z. siniestri'' * ''Z. viridis'' ;
La Pedrera de Rúbies Formation The La Pedrera de Rúbies Formation is an Early Cretaceous ( late Berriasian to early Barremian geologic formation in Catalonia, Spain. The formation crops out in the area of the Montsec in the Organyà Basin. At the La Pedrera de Meià localit ...
, Las Hoyas, Spain * ''Z. martinclosas'' * ''Z. oculata'' ; Ulan-Argalant Formation, Mongolia * ''Z. mongolicus'' ;
Weald Clay Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of ro ...
, United Kingdom * ''Z. angliae'' * ''Z. floodpagei'' * ''Z. tuberculata'' * ''Z. simpsoni''


Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
(125.0 to ~113.0 Ma)

; Argun Formation, Russia * ''Z. semen'' * ''Z. sibirica'' * ''Z. sinitzae'' ; Dzun-Bain Formation, Mongolia * ''Z. dundulaensis'' * ''Z. elegans'' ;
Jehol Group The Jehol Biota includes all the living organisms – the ecosystem – of northeastern China between 133 and 120 million years ago. This is the Lower Cretaceous ecosystem which left fossils in the Yixian Formation and Jiufotang Formation. These ...
, China * ''Z. valida'' ; Khasurty Formation, Russia * ''Z. khasurtyiensis'' ;
Laiyang Formation Laiyang city () is a county-level city within Yantai bordering Qingdao, located in the middle of the Shandong Peninsula, in Shandong province, China. The majority (70%) of its population are farmers and it is famous for producing the Laiyang pe ...
, China * ''Z. laiyangensis'' * ''Z. ludongensis'' * ''Z. tuanwangensis'' ; Shahai Formation, China * ''Z. kezuoensis'' ;
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...
, China * ''Z. aliena'' * ''Z. epicharis'' * ''Z. eumeura'' * ''Z. liui'' * ''Z. miniscula'' * ''Z. porrecta'' * ''Z. protensa'' * ''Z. psilata'' * ''Z. rudis'' * ''Z. stabilis'' ;
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous period. It is Aptian in age and consists of Sandstone, sandstones, Siltstone, siltstones, Marl, marls and bituminous Shale, shales, deposi ...
, Russia * ''Z. caudatus'' * ''Z. excellens'' * ''Z. vitimensis''


Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...
(93.9 to 89.8 Ma)

; Emanra Formation, Russia * ''Z. khetanensis'' ;Kzyl-Zhar Locality, Kazakhstan * ''Z. caduca''


References

Ommatidae Prehistoric beetle genera {{Ommatidae-stub